500 ‘Genetic Zipper’ Technology-Based Oligonucleotide Pesticides Generated in 15 Minutes Using DNAInsector Algorithm (dnainsector.com): Fast and Potent Tool for Pest Control

Vol Oberemok; Kate Laikova; Oksana Andreeva; Anastasia Dmitrienko; Anastasia Shapovalova; Nikita Stolyarchuk; Nikita Gal'chinsky

doi:10.20944/preprints202509.1765.v2

Submitted:

30 January 2026

Posted:

02 February 2026

You are already at the latest version

Abstract

The article provides basic knowledge on how to calculate selectivity of oligonucleotide pesticides when ‘genetic zipper’ (‘GZ’) technology is used to provide safety for non-target organisms. Also here we represent 500 ‘GZ’ technology-based oligonucleotide pesticides generated in 15 minutes using DNAInsector algorithm (dnainsector.com) to show that ‘GZ’ technology is amazing approach transferring us to a new oligo era of development of potent and selective oligonucleotide pesticides based on new principles prompted by nature.

Keywords:

DNA insecticides

;

oligonucleotide pesticides

;

olinscides

;

rRNA

Subject:

Biology and Life Sciences - Biology and Biotechnology

Introduction

Today, in plant protection, there is an understanding that it is necessary to create genus- and species-specific pesticides that could solve problems with globally important pests being safe for non-target organisms [1]. Oligonucleotide pesticides based on ‘genetic zipper’ (‘GZ’) technology have an easily adaptable structure and a selective DNA containment mechanism of action, which opens up the possibility of effective and well-tailored pest control with minimal side effects [2,3]. Oligonucleotide pesticides can be designed very quickly using DNAInsector program (available at dnainsector.com) or manually, based on pre-rRNA and mature rRNA sequences retrieved from the GenBank database [4]. In practical terms, this means that individuals with basic sequence knowledge can quickly design an oligonucleotide pesticide complementary to the pre-rRNA or mature rRNA of a susceptible to this approach pest with a high probability of success. In this article we represent 500 ‘GZ’ technology-based oligonucleotide pesticides generated in 15 minutes using DNAInsector algorithm (dnainsector.com) to show that ‘GZ’ technology is amazing approach transferring us to a new oligo era of development of potent and selective oligonucleotide pesticides based on new principles prompted by nature (Table 1). Already today, the ‘GZ’ technology is potentially capable of controlling 20% of all invertebrate pests with a simple and flexible algorithm [3]. However, to ensure species specificity, it is essential to compare homologous target sites of pre-rRNA and mature rRNA in non-target organisms to prevent off-target effects [3,4].

How to Calculate Selectivity of Oligonucleotide Pesticides?

A very important preoperative recommendation is how to calculate the safety of a specific oligonucleotide pesticide in an ecosystem for a range of non-target organisms [4]. To do this, select a target rRNA region (for example, 500 nt long fragment of 28S rRNA) of a pest and corresponding regions of non-target organisms. This requires knowing, after DNA sequencing, the target rRNA sequence (in our case it is 500 nt long fragment of 28S rRNA) of all invertebrates of a given ecosystem (other animals and plants will not be susceptible to ‘GZ’ technology due to physiological barriers). Next, align target 500 nt long fragment of 28S rRNA of all invertebrates of a given ecosystem, select the most differing part of the sequence that belongs to pest (it can be ~100 nt long sequence out of 500 nt long fragment of 28S rRNA), and then upload it to the DNAInsector web tool. Generate unique oligonucleotide pesticide 11 nt long and check one more time, adjust manually if required, that it is not complementary elsewhere to the target 500 nt long fragment of 28S rRNA of non-target invertebrates. Mathematically, with a high probability (>99.9%), this oligonucleotide pesticide will also not be complementary to other rRNAs of non-target invertebrates in a given ecosystem. Therefore, information on total rRNA (which includes 5S, 5.8S, 12S, 16S, 18S, 28S, ETS and ITS regions and comprising ca. 10,000 nt in each non-target invertebrate) is not required, what significantly simplifies the selection of potent and selective oligonucleotide pesticides [4].

Beginning from 2008, in our research work we found that pre-rRNA and mature rRNA is a convenient target for oligonucleotide pesticides, while mRNA, due to much lower concentration, will be much less susceptible to them, even if oligonucleotide pesticides will possess perfect complementarity to it. Pest rRNA comprises 80-85% of all RNA in the cell [5] and its use as a target for ‘GZ’ technology helps making this approach very efficient and selective at the same time [2,4]. Thousands of different mRNAs make up only 5% of all RNA and use of mature rRNA and pre-rRNA for targeting substantially increases signal-to-noise ratio, ca. 10⁵:1 (rRNA vs. random mRNA) [6].

Importantly, for ‘GZ’ technology it would be easy to create algorithms (plans) of designing potent and selective oligonucleotide pesticides for other cases, such as when an ecosystem contains two major pests and a dozen non-target organisms or when one species is pest and closely related species is beneficial, and many other scenarios of pest control [4]. The algorithm of ‘GZ’ technology is very simple and efficient, fundamentally different from all modern approaches to plant protection, including RNAi and CRISPR/Cas [3]. In a sense, ‘GZ’ technology is an oligo universe for the scientific exploration by a biologist or a plant protection specialist. And if they enter this oligo universe, it will enrich their capabilities.

Author Contributions

Conceptualization, V.O.; methodology, K.L., O.A., A.D., A.S., N.S. and N.G.; software, N.G.; validation, V.O.; formal analysis, V.O. and N.G.; investigation, V.O., K.L., O.A., A.D., A.S., N.S. and N.G.; resources, V.O.; data curation, V.O.; writing—original draft preparation, V.O. and N.G.; writing—review and editing, V.O. and N.G.; project administration, V.O.; funding acquisition, V.O. All authors have read and agreed to the published version of the manuscript.

Funding

The research obtained funding from the Russian Science Foundation No. 25-16-20070, https://rscf.ru/project/25-16-20070/ (accessed on 30 January 2026).

Data Availability Statement

The original contributions presented in this study are included in the article. Further inquiries can be directed to the corresponding authors.

Acknowledgments

We thank our many colleagues, too numerous to name, for the technical advances and lively discussions that have prompted us to write this article. We apologize to the many colleagues whose work has not been cited. Research work was carried out at the Molecular Genetics and Biotechnologies Lab created within the framework of a state assignment V.I. Vernadsky Crimean Federal University for 2026 and the planning period of 2024–2026 No. FZEG-2024–0001. We are very much indebted to all anonymous reviewers and our colleagues from the Lab for DNA technologies, PCR analysis, and the creation of DNA insecticides (V.I. Vernadsky Crimean Federal University, Institute of Biochemical Technologies, Ecology and Pharmacy, Department of General Biology and Genetics), and OLINSCIDE BIOTECH LLC. for the valuable comments on our manuscript.

Conflicts of Interest

The authors declare no conflicts of interest.

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Table 1. 500 ‘genetic zipper’ technology-based oligonucleotide pesticides generated in 15 minutes using DNAInsector algorithm (dnainsector.com).

№	Reference	Latin name	GenBank ID	Target rRNA	Sequence of oligonucleotide pesticide (5’..3’)
APHIDOIDEA
	https://doi.org/10.1007/s10886-012-0121-y	Brachycaudus helichrysi	JX965980.1	12S	ACTAAAATACC
	2. https://doi.org/10.1093/jee/toae041	Myzus cerasi	KX631445.1	16S	CATACAAGTCC
	3. http://dx.doi.org/10.3733/ca.v054n06p26	Aphis gossypii	HG810150.1	16S	TAAATATTAGG
	4. https://doi.org/10.1042/bj0660289	Eucallipterus tiliae	KX631489.1	16S	CATTCTAGTCC
	5. https://doi.org/10.1093/jee/toy157	Metopolophium dirhodum	XR_009664640.1	5.8S	TTCATCGATCC
	6. http://dx.doi.org/10.18801/jstei.080119.58	Rhopalosiphum maidis	XR_003406101.1	5S	ACTAACTACGC
	7. http://dx.doi.org/10.1603/0046-225X-34.4.938	Hysteroneura setariae	MT116412.1	12S	TAAAATACCGC
	8. https://doi.org/10.1073/pnas.0703535104	Megoura viciae	EU071373.1	ITS2	TCACAACTTGG
	9. https://doi.org/10.1201/9781003169239	Myzus persicae	HG810161.1	16S	AATTAGTTTGC
	10. http://dx.doi.org/10.1515/ausae-2016-0007	Macrosiphum rosae	JX966002.1	12S	TTATAATAGGG
	11. http://dx.doi.org/10.4289/0013-8797.119.1.90	Aphis ruborum	JX965974.1	12S	ATTAATCGTGG
	12. https://doi.org/10.1016/j.asd.2016.01.005	Eriosoma lanigerum	HG810182.1	16S	TTAATCTTAGG
	13. https://doi.org/10.1016/j.plantsci.2014.01.010	Acyrthosiphon pisum	KT199046.1	18S	AAGTAACTTGG
	14. https://doi.org/10.1016/j.micpath.2024.107015	Aphis craccivora	JX965963.1	12S	TAATAAACAGG
	15. https://doi.org/10.1371/journal.pone.0223616	Aphis fabae	KC897347.1	16S	ATTACTCACCC
	16. https://doi.org/10.1017/s0007485320000097	Brevicoryne brassicae	HG810153.1	16S	TAAGTATTAGG
	17. https://doi.org/10.1007/s42976-023-00370-w	Macrosiphum euphorbiae	KX631452.1	16S	CAAAATACTGC
	18. https://doi.org/10.1371/journal.pone.0284888	Sitobion avenae	AY745779.1	16S	GCTACCTTTGC
	19. https://doi.org/10.1017/s0007485323000603	Aphis spiraecola	KC897427.1	16S	TATTACTCACC
	20. https://doi.org/10.1371/journal.pone.0017524	Aphis nerii	KC897401.1	16S	CCTCTTTTGGG
	21. https://doi.org/10.3897/zookeys.319.4315	Brachycaudus cardui	JX965979.1	12S	ATTTATAAACC
	22. https://doi.org/10.3897/zookeys.319.4315	Hyperomyzus pallidus	JX965997.1	12S	GTATCTAATCC
	23. https://doi.org/10.3897/zookeys.319.4318	Idiopterus nephrelepidis	KX631469.1	16S	ATAATTTAAGC
	24. https://doi.org/10.1007/s10886-018-0980-y	Phorodon humuli	JX966008.1	12S	TCTATTCCTGG
	25. https://doi.org/10.1673/031.012.14301	Pterochloroides persicae	PP601230.1	12S	AAATGTACTGG
	26. https://doi.org/10.3390/insects14110862	Sipha maydis	OQ401327.1	18S	TTGAAAGATGC
	27. https://doi.org/10.3390/insects15100816	Aphis glycines	KC897361.1	16S	CCCTCTTTTGG
	28. https://doi.org/10.3390/insects12080744	Aphis nasturtii	KC897399.1	16S	AGTTATCCCCC
	29. https://doi.org/10.3897/zookeys.319.4318	Aulacorthum solani	HG810152.1	16S	TAAATCTTAGG
	30. https://doi.org/10.1038/s41598-018-27933-w	Brachycaudus persicae	JX965981.1	12S	TATTGTAACCC
	31. https://doi.org/10.1155/2013/838145	Capitophorus elaeagni	KX631436.1	16S	AAAATTTAAGC
	32. https://doi.org/10.3390/ijms23137195	Ceratovacuna lanigera	AB035877.1	16S	GATATAAAAGG
	33. https://doi.org/10.1128/JB.186.19.6626-6633.2004	Cinara cedri	CABPRJ010000001.1	5S	TTAGTACTTGG
	34. https://doi.org/10.1371/journal.pone.0012053	Diuraphis muehlei	NC_022727.1	12S	ACATATTGCCC
	35. https://doi.org/10.1186/1471-2148-12-106	Geoica utricularia	PP486353.1	16S	TTACTTTAGGG
	36. https://doi.org/10.1038/s41598-019-46441-z	Lachnus roboris	PP601231.1	12S	TACATATTGCC
	37. https://doi.org/10.1186/s40064-015-1263-5	Pemphigus bursarius	JX966006.1	12S	TTTAAAGAACC
	38. https://doi.org/10.1111/mec.14506	Prociphilus fraxinifolii	AF275220.1	16S	GAAATTAAAGG
	39. https://doi.org/10.3390/insects14070639	Rhopalosiphum rufiabdominale	EU358897.1	16S	ATACTAAAACC
	40. https://doi.org/10.3390/insects11080460	Tuberolachnus salignus	AF275246.1	16S	TTAATTAACCC
	41. https://doi.org/10.1016/j.ygeno.2022.110472	Toxoptera aurantii	JX966017.1	12S	TTTATAATAGG
	42. https://doi.org/10.3390/v16060919	Cavariella aegopodii	HG810155.1	16S	ATCTTAGGTGC
	43. https://doi.org/10.1016/j.jinsphys.2015.10.007	Pemphigus betae	U27824.1	18S	ATTTAATGAGC
	44. https://doi.org/10.1016/0042-6822(68)90213-4	Hyperomyzus lactucae	KX631467.1	16S	AATTATTATGC
	45. https://doi.org/10.1093/cz/zox004	Cerataphis brasiliensis	AB301599.1	16S	TTTAATTAACC
	46. https://doi.org/10.3390/insects10040109	Hyalopterus amygdali	OK641613.1	12S	GTACATATTGC
	47. https://doi.org/10.3390/insects12020173	Hyalopterus pruni	KC753377.1	ITS1	TTTTGCCTTGC
	48. https://doi.org/10.1673/031.013.8301	Myzus ascalonicus	Y07924.1	ITS1	GTGATCCTTCC
	49. https://doi.org/10.1111/j.1570-7458.2011.01213.x	Myzus varians	KX631439.1	16S	ATACAAGTCCC
	50. https://doi.org/10.1093/jis/5.1.13	Schizolachnus pineti	KP637088.1	ITS2	TTTGCCATCGC
	51. https://doi.org/10.3390/genes12121966	Aphis hederae	JX965968.1	12S	TTATTGTAACC
	52. https://doi.org/10.1093/jisesa/ieu033	Aphis farinosa	KC897352.1	16S	TAGTTATCCCC
	53. https://doi.org/10.1098/rspb.2008.0679	Brachycaudus tragopogonis	OZ060884.1	16S	GTTACTTTAGG
	54. https://doi.org/10.1038/s41598-019-46441-z	Cavariella theobaldi	OZ060905.1	12S	AATGAAATTGG
	55. https://doi.org/10.3390/insects10100314	Myzus certus	Y07926.1	ITS1	TCTAATGATCC
	56. https://doi.org/10.3390/insects11050282	Pemphigus spyrothecae	JX966007.1	12S	TAAAAAACAGG
	57. https://doi.org/10.1098/rspb.2004.3033	Sitobion fragariae	JX966013.1	12S	ATTTATTAACC
	58. https://doi.org/10.3390/plants11030244	Tetraneura ulmi	OZ060885.1	16S	AATAAAATTGG
	59. https://doi.org/10.3389/fpls.2024.1441145	Amphorophora idaei	X78807.1	ITS1	TTTCTACGGCC
	60. https://doi.org/10.1371/journal.pone.0184080	Nasonovia ribisnigri	KX631447.1	16S	CTTATCGTCCC
	61. https://doi.org/10.3389/fmicb.2022.961349	Aphis illinoisensis	KC897378.1	16S	GTAGTTATCCC
	62. https://doi.org/10.2903/j.efsa.2024.8657	Anoecia corni	OZ060897.1	16S	TTAATTGTAGG
	63. https://doi.org/10.3390/insects12020173	Aphis sambuci	KC897412.1	16S	CTCTTTTGGGC
	64. https://doi.org/10.3390/insects12020173	Chaetosiphon tetrarhodum	KX631472.1	16S	GTTTTATAGGG
	65. https://doi.org/10.1007/s00248-021-01937-y	Myzus ornatus	Y07927.1	ITS1	GTATAACCTGC
	66. https://doi.org/10.3390/insects15090633	Rhopalosiphum nymphaeae	JX966010.1	12S	TAATGTAACCC
	67. https://doi.org/10.3897/zookeys.221.3541	Sipha elegans	HM142963.1	18S	GTCCTCATTCC
	68. https://doi.org/10.1094/pdis-07-15-0805-re	Uroleucon sonchi	KX631442.1	16S	AGTTTTATAGG
	69. https://doi.org/10.1007/s00248-016-0892-8	Aphis citricidus	MG988610.1	18S	TTAATATACGC
	70. https://doi.org/10.1603/ec13263	Sipha flava	AY745776.1	16S	TTTATTTAACC
	71. https://doi.org/10.2903/j.efsa.2018.5103	Toxoptera citricida	EU358901.1	16S	TTTAATCAACC
PHYLLOXEROIDEA
	72. https://doi.org/10.1371/journal.pone.0251663	Adelges laricis	AF275215.1	12S	AATAAAATACC
	73. https://doi.org/10.1016/j.foreco.2013.05.047	Adelges tsugae	KT199045.1	18S	CACCTTAATGC
	74. https://doi.org/10.1038/s41598-020-77928-9	Daktulosphaira vitifoliae	MK685299.1	16S	GAATTTAAAGG
ALEYRODIDAE
	75. https://doi.org/10.1111/epp.12760	Aleurocanthus camelliae	OQ180914.1	16S	TTTACTGCAGC
	76. https://doi.org/10.3390/insects11010042	Aleurocanthus spiniferus	OQ180915.1	16S	TAATTTACTGC
	77. https://doi.org/10.1080/21658005.2013.795042	Aleurochiton aceris	AY521267.1	16S	TAAAATTTTGG
	78. https://doi.org/10.61310/mndjstea.1015.23	Aleurodicus dispersus	JQ305695.1	16S	TTTCATTGAGC
	79. https://doi.org/10.1016/S0305-0491(99)00054-1	Aleurodicus dugesii	U06474.1	18S	TTAGAACTAGG
	80. https://doi.org/10.3956/2010-05.1	Aleuroplatus coronata	EU441164.1	16S	TGTTAAACAGG
	81. https://doi.org/10.3956/2010-05.1	Aleuroplatus gelatinosus	EU441163.1	16S	AAATTTAAGCC
	82. https://doi.org/10.1079/cabicompendium.4538	Aleurothrixus floccosus	HG810135.1	16S	TCTGTTCGACC
	83. https://doi.org/10.1093/jisesa/ieu136	Aleurotrachelus camelliae	OQ180920.1	16S	TTATTACAACC
	84. https://doi.org/10.1007/s41348-020-00319-9	Aleyrodes proletella	MW644572.1	16S	ATTTACTGCGG
	85. https://doi.org/10.1007/978-3-030-28683-5_8	Bemisia afer	GQ867747.1	16S	AATCATTGAGC
	86. https://doi.org/10.1093/jee/99.3.691	Bemisia argentifolii	AY521257.1	12S	TATCAGATTGC
	87. https://doi.org/10.1007/s13205-021-02831-7	Bemisia breyniae	GQ867745.1	16S	TGTCATTGAGC
	88. https://doi.org/10.1038/s41598-017-00528-7	Bemisia emiliae	GQ867744.1	16S	ATTCATTGAGC
	89. https://doi.org/10.1007/s13205-021-02831-7	Bemisia euphorbiae	GQ867743.1	16S	ATTATGCTACC
	90. https://doi.org/10.1007/s10340-020-01210-0	Bemisia tabaci	AF110722.3	16S	ATTTATTACGC
	91. https://doi.org/10.1007/s12600-023-01059-w	Paraleyrodes bondari	GQ867760.1	16S	TCTCATTGAGC
	92. https://doi.org/10.1093/jisesa/ieu136	Pealius mori	HG810145.1	16S	TTGGATTAAGC
	93. https://doi.org/10.5656/KSAE.2012.04.1.83	Pealius rhododendri	GQ867752.1	16S	GTTCATTGAGC
	94. https://doi.org/10.1673/031.010.12001	Siphoninus phillyreae	Z15053.1	18S	GTTAGCTTTGG
	95. https://doi.org/10.5958/0974-8172.2018.00158.X	Tetraleurodes acaciae	ON311124.1	12S	CCTATAAAAGG
	96. https://doi.org/10.1111/j.1365-2311.2004.00586.x	Tetraleurodes mori	AY521263.1	12S	CACTCTTAAGC
	97. https://doi.org/10.1038/s41598-024-84958-0	Trialeurodes ricini	HG810146.1	16S	ATAAGATTAGG
	98. https://doi.org/10.1093/gbe/evaa216	Aleurolobus barodensis	PQ863114.1	16S	GTTAAAATCGG
COCCOIDEA
	99. https://doi.org/10.3390/insects11070399	Acanthococcus lagerstroemiae	KJ869290.1	28S	TTTCATTGCGC
	100. https://doi.org/10.1093/jisesa/ieae032	Acutaspis scutiformis	KY219127.1	28S	TTCATTACGCC
	101. https://doi.org/10.1111/syen.12033	Affirmaspis cederbergensis	MH934019.1	28S	CCATCTTCCCC
	102. https://doi.org/10.12681/eh.13916	Anapulvinaria pistaciae	OR074914.1	5.8S	GTTTGTACAGC
	103. https://doi.org/10.11646/zootaxa.4117.1.4	Ancepaspis edentata	KY220051.1	28S	ACCTACTGTCC
	104. https://doi.org/10.21608/sjas.2024.322627.1461	Aonidiella pini	MK886648.1	28S	ATCAAACAACC
	105. https://doi.org/10.3897/zookeys.867.34937	Aspidaspis florenciae	KY219598.1	28S	ACCATCTTCCC
	106. https://doi.org/10.1079/cabicompendium.7490	Aspidiella hartii	KY219906.1	28S	TCCTGAATACC
	107. https://doi.org/10.1079/cabicompendium.7506	Aspidiella sacchari	DQ145368.2	28S	TCCCGTTTACC
	108. https://doi.org/10.3897/zookeys.1047.68409	Aspidiotus fularum	MH934073.1	28S	TTTCATTACGC
	109. https://doi.org/10.3897/zookeys.1174.105851	Aulacaspis difficilis	DQ145298.2	28S	ATATCAAACGG
	110. https://doi.org/10.3897/zookeys.1174.105851	Aulacaspis distylii	DQ145299.2	28S	AGTCTTTCGCC
	111. https://doi.org/10.1111/epp.12778	Aulacaspis rosae	KY219387.1	28S	GCTTACTGTCC
	112. https://doi.org/10.1016/S1226-8615(08)60379-9	Aulacaspis spinosa	DQ145367.2	28S	GCAATTCCTCC
	113. https://doi.org/10.11646/zootaxa.4272.1.6	Austrolecanium cryptocaryae	KY816393.1	18S	TTTAATGAGCC
	114. https://doi.org/10.11646/zootaxa.4272.1.6	Austrolecanium sassafras	KY816397.1	18S	CAAGTCTTTGC
	115. https://doi.org/10.11646/zootaxa.4508.1.6	Austrolichtensia hakearum	MH844461.1	18S	TTTCACCGTGC
	116. https://doi.org/10.61310/mjst.v23i1.2370	Ceroplastes floridensis	JQ795604.1	28S	TCCTGAATTCC
	117. https://doi.org/10.1007/s13744-016-0480-0	Ceroplastes glomeratus	KX670822.1	28S	TTCAACTTTCC
	118. https://doi.org/10.11646/zootaxa.4701.6.2	Ceroplastes kunmingensis	MT316993.1	28S	CTTCATCCTGG
	119. https://doi.org/10.11646/zootaxa.4701.6.2	Ceroplastes murrayi	MT316994.1	28S	ACTTTCATTGC
	120. https://doi.org/10.1079/pwkb.species.12351 120. https://doi.org/10.2903/j.efsa.2024.8888	Ceroplastes rubens	MT317009.1	28S	TTCATTGCGCC
	121. https://doi.org/10.11609/jott.7419.14.2.20606-20614	Ceroplastes rusci	PV762166.1	28S	CATCTTCCCCC
	122. https://doi.org/10.1079/cabicompendium.12353	Ceroplastes sinensis	KY085826.1	28S	GTCCGTTTACC
	123. https://doi.org/10.1079/cabicompendium.120372	Ceroplastes stellifer	MK533217.1	28S	TCCGTTTACCC
	124. https://doi.org/10.1111/j.1095-8312.2011.01716.x	Chionaspis americana	KY220045.1	28S	TTGAATTCCGC
	125. https://doi.org/10.1111/epp.12287	Chionaspis etrusca	DQ145397.2	28S	GATCGATTTGC
	126. https://doi.org/10.1590/S1519-566X2010000300013	Coccus alpinus	JX499976.1	28S	AAATTCATCGC
	127. https://doi.org/10.1079/cabicompendium.14663	Coccus celatus	MT317016.1	28S	CCATCTTTCGG
	128. https://doi.org/10.3897/zookeys.734.22774	Coccus ficicola	MK533218.1	28S	TTACTCCTCGG
	129. https://doi.org/10.3897/zookeys.244.4045	Coccus formicarii	MZ782004.1	28S	CCTCGATTACC
	130. https://doi.org/10.1017/S0007485300010919	Coccus longulus	MT317024.1	28S	TGACTTCATCC
	131. https://doi.org/10.11646/zootaxa.4508.1.6	Cryptes baccatus	MZ782005.1	28S	TATATCGTCGG
	132. https://doi.org/10.11646/zootaxa.4508.1.6	Cryptes utzoni	MH886632.1	28S	CATAGTTCACC
	133. https://doi.org/10.1080/23802359.2021.1906175	Didesmococcus koreanus	MH844459.1	18S	CATGTATTAGC
	134. https://doi.org/10.1093/gigascience/giz113	Ericerus pela	KX380986.1	18S	CCAATTGATCC
	135. https://doi.org/10.1603/0022-0493-98.4.1202	Eulecanium cerasorum	MK533231.1	28S	TAGTCTTTCGC
	136. https://doi.org/10.1007/s10530-018-1749-5	Nipponaclerda biwakoensis	MZ824101.1	28S	CCACATTTCCC
	137. https://doi.org/10.2307/3496360	Planchonia stentae	JX500033.1	28S	GTTTACACAGC
	138. https://doi.org/10.21506/j.ponte.2020.1.17	Dactylopius opuntiae	MN310167.1	18S	TTTACTGAGCC
	139. https://doi.org/10.53555/jaz.v44iS5.2877	Dactylopius ceylonicus	MN310174.1	18S	ATTTACTGAGC
	140. https://doi.org/10.17660/ActaHortic.2022.1343.73	Dactylopius coccus	KJ701995.1	18S	ACAATCTAAGG
	141. https://doi.org/10.5897/IJBC2018.1204	Dactylopius tomentosus	MN310135.1	18S	ACCCTGATTCC
	142. https://doi.org/10.5281/zenodo.185547	Abgrallaspis aguacatae	GQ478401.1	28S	TACCAACATGC
	143. https://doi.org/10.1080/095831500750016389	Abgrallaspis cyanophylli	JQ651047.1	18S	ACTATCTAAGG
	144. https://doi.org/10.2298/PIF1602069G	Aonidia lauri	KY219724.1	28S	TAAAGTAACGC
	145. https://doi.org/10.28955/alinterizbd.741562	Icerya purchasi	AY426078.1	18S	AACTCTATCGG
	146. https://doi.org/10.2903/j.efsa.2014.3929	Aonidiella citrina	GQ325448.1	28S	CCCTGAATTCC
	147. https://doi.org/10.1088/1755-1315/1252/1/012007	Aonidiella orientalis	KY219919.1	28S	ATAACGTTCGC
	148. https://doi.org/10.13140/RG.2.2.26296.67846	Aspidiotus destructor	JQ650955.1	18S	GTAATTTGCGC
	149. https://doi.org/10.1603/AN10060	Aspidiotus nerii	JQ650991.1	18S	TACCCGTTACC
	150. https://doi.org/10.2903/j.efsa.2022.7307	Aulacaspis tubercularis	JQ651055.1	18S	TCACTACCTCC
	151. https://doi.org/10.1111/1442-1984.12505	Aulacaspis yasumatsui	KX091314.1	28S	TTACTGTCCGC
	152. https://doi.org/10.1093/jee/98.5.1603	Chionaspis heterophyllae	GU349114.1	28S	ATCGCAATTCC
	153. https://doi.org/10.1603/0046-225X-29.6.1305	Chionaspis pinifoliae	MH032863.1	28S	TCTTGAATTCC
	154. https://doi.org/10.4081/jear.2010.147	Chionaspis wistariae	GU349092.1	28S	TTACGCCTTCC
	155. https://doi.org/10.1080/00779962.2013.795646	Diaspidiotus perniciosus	JQ650906.1	18S	TCGTCACTACC
	156. https://doi.org/10.12741/ebrasilis.v13.e0880	Pinnaspis strachani	GQ284590.1	18S	ATGATCCTTCC
	157. https://doi.org/10.5296/jas.v8i4.17519	Pseudaulacaspis pentagona	HG810175.1	16S	AAATTATTAGG
	158. https://doi.org/10.3390/insects11110745	Aspidiotus rigidus	MG940953.1	ITS2	GATACTTCAGC
	159. https://doi.org/10.5154/r.inagbi.2021.01.005	Hemiberlesia lataniae	GQ284610.1	ITS2	TCTTTTGACGG
	160. https://doi.org/10.4039/tce.2020.4	Eriococcus spurius	AY795540.1	18S	TTTCAGAACGC
	161. https://doi.org/10.1016/j.biocontrol.2006.04.009	Cryptococcus fagisuga	DQ125262.1	18S	AACTCAATAGG
	162. https://doi.org/10.1515/biol-2021-0066	Marchalina hellenica	KT199030.1	18S	GTTCTTTCCGC
	163. https://doi.org/10.1111/jbi.13702	Matsucoccus macrocicatrices	KF053091.1	18S	TTCAGAATCGC
	164. https://doi.org/10.1653/0015-4040(2008)91[335:ANEPFF]2.0.CO;2	Crypticerya genistae	EU087718.1	18S	CATTTGAGTCC
	165. https://doi.org/10.1016/j.jcz.2017.12.004	Drosicha corpulenta	KT199038.1	18S	TTGATCCTCGC
	166. http://dx.doi.org/10.17582/journal.pjz/2019.51.5.1815.1822	Drosicha mangiferae	AB523733.1	18S	TTAAAGTGTGC
	167. https://doi.org/10.2903/j.efsa.2023.7739	Icerya aegyptiaca	EU087752.1	18S	ATTTGAGTCCC
	168. https://doi.org/10.1093/jipm/pmae018	Icerya seychellarum	EU087747.1	18S	CTTAGACATGC
	169. https://doi.org/10.1007/s42690-020-00382-7	Antonina graminis	AY426047.1	18S	TTCCTAAATGC
	170. https://doi.org/10.9734/ijpss/2024/v36i44477	Dysmicoccus brevipes	AY426037.1	18S	AATCTTTGCGC
	171. https://doi.org/10.18520/cs/v128/i9/946-950	Dysmicoccus neobrevipes	KU891794.1	18S	CAAATCTTTGC
	172. https://doi.org/10.1093/jee/tow124	Ferrisia gilli	AY426067.1	18S	TCAATTGATCC
	173. https://doi.org/10.3390/insects14090720	Planococcus citri	XM_065345395.1	5S	GTGTATTCAGC
	174. https://doi.org/10.1093/jee/toz125	Pseudococcus longispinus	HG810200.1	16S	AGATTATTACC
	175. https://doi.org/10.3390/insects12040343	Pseudococcus calceolariae	MG866178.1	28S	TTCTGAATTCC
	176. https://doi.org/10.1371/journal.pone.0157965	Pseudococcus comstocki	KU499441.1	28S	AATATCGAACC
	177. https://doi.org/10.1002/ps.6565	Phenacoccus solenopsis	HG810195.1	16S	AGTTTATTACC
	178. https://doi.org/10.3390/insects14040345	Pulvinaria hydrangeae	MT317058.1	28S	TTCGATTACCC
	179. https://doi.org/10.3390/v13102081	Pulvinaria vitis	OR678175.1	28S	CTTCGATTACC
	180. https://doi.org/10.3390/v14122679	Parthenolecanium corni	MK533241.1	28S	TCGTTTACCCC
	181. https://doi.org/10.1093/jee/tox170	Parthenolecanium persicae	MK533243.1	28S	TTCGTTTACCC
	182. https://doi.org/10.1093/ee/nvaa141	Planococcus ficus	LC583719.1	18S	TCTAAGAACCC
	183. https://doi.org/10.1590/1519-6984.237273	Dysmicoccus boninsis	MW542087.1	28S	TTTCCAAGCCC
	184. https://doi.org/10.2903/j.efsa.2018.5187	Unaspis citri	KY219855.1	28S	AATATCAAAGG
	185. https://doi.org/10.3390/plants13070952	Unaspis yanonensis	KY219678.1	28S	TACTGTCCACC
	186. https://doi.org/10.3897/zookeys.309.5318	Lepidosaphes beckii	DQ145352.2	28S	ATTTCGTCGCC
	187. https://doi.org/10.3897/zookeys.309.5318	Lepidosaphes gloverii	DQ145353.2	28S	TGAATTTCGCC
	188. https://doi.org/10.2903/j.efsa.2022.7024	Maconellicoccus hirsutus	PX279479.1	28S	AACTTTAGCGC
	189. https://doi.org/10.3897/zookeys.779.27437	Parlatoria pergandii	DQ145372.2	28S	CACCATCTTCC
	190. https://doi.org/10.3390/insects16101070	Parlatoria ziziphi	KY219837.1	28S	TGTTAGACTCC
	191. https://doi.org/10.3897/zookeys.779.27437	Parlatoria oleae	KY219149.1	28S	TACCAATGACC
	192. https://doi.org/10.2903/j.efsa.2017.4926	Phenacoccus aceris	KY940032.1	28S	ATTCTGCTGCC
	193. https://doi.org/10.3832/ifor0703-006	Physokermes piceae	OQ657453.1	28S	TTCATCCTGGC
	194. https://doi.org/10.11646/zootaxa.5195.1.6	Protopulvinaria pyriformis	PV153706.1	28S	TACTTGTCCGC
	195. https://doi.org/10.1017/S0007485300015558	Rastrococcus invadens	PX279482.1	28S	CTCTTAAGAGC
	196. https://doi.org/10.21608/eajbsa.2009.15438	Saissetia coffeae	HG810170.1	16S	AGATTATGACC
	197. https://doi.org/10.3897/zookeys.873.36661	Saissetia oleae	MT276989.1	18S	TAGAACCTACC
	198. https://doi.org/10.1007/s12600-018-0678-2	Kermes echinatus	JX436139.1	28S	TTCTAACCGCC
	199. https://doi.org/10.2903/j.efsa.2024.8666	Eulecanium giganteum	KJ908303.1	28S	GTTTAGTAAGC
	200. https://doi.org/10.1007/s12600-024-01175-1	Megapulvinaria maxima	KP189631.1	28S	AATAAAAATCC
	201. https://doi.org/10.2903/j.efsa.2023.7846	Milviscutulus mangiferae	JX645355.1	28S	TTCCATTCGCC
	202. https://doi.org/10.1653/024.097.0302	Parasaissetia nigra	KY933367.1	28S	TCTTTCGCCCC
	203. https://doi.org/10.55446/IJE.2021.56	Prococcus acutissimus	KP189525.1	28S	GTTCGCTATCC
	204. https://doi.org/10.1080/09064710.2017.1338745	Pulvinaria aurantii	MT317050.1	28S	ACTACTACCCC
	205. https://doi.org/10.2903/j.efsa.2022.7526	Pulvinaria psidii	JQ651036.1	18S	CTTAATTCGGC
	206. https://doi.org/10.3733/hilg.v53n02p027	Pulvinariella mesembryanthemi	JQ651137.1	18S	ATCTCTACTGG
	207. https://doi.org/10.1111/epp.13089	Saissetia miranda	JX866682.1	18S	ACTATCTCTGC
	208. https://doi.org/10.2903/j.efsa.2023.8000	Takahashia japonica	MK533207.1	18S	ATTTGGCATGC
	209. https://doi.org/10.24425/jwld.2025.154265	Aonidiella aurantii	U06475.1	18S	ATTTAAACGGC
	210. https://doi.org/10.5281/zenodo.10930744	Eriococcus buxi	AY795513.1	18S	GTAATTTACGC
	211. https://doi.org/10.1111/j.1365-3113.1958.tb00403.x 211. https://doi.org/10.36305/2712-7788-2022-4-165-7-16	Ovaticoccus agavium	KT199025.1	18S	CTAATAAGAGC
	212. https://doi.org/10.1093/jee/toad184	Thysanococcus pandani	DQ145391.2	28S	TTCCAATGACC
	213. https://doi.org/10.18698/2542-1468-2022-2-31-35	Kermes quercus	JX436143.1	28S	ATTCTAACCGC
	214. https://doi.org/10.2903/j.efsa.2019.5672	Eumargarodes laingi	KT199035.1	18S	GTACTCATTGC
	215. https://doi.org/10.1603/en13004	Eurhizococcus brasiliensis	OK085528.1	18S	AGTAATTGTGG
	216. https://doi.org/10.5281/zenodo.4565418	Matsucoccus alabamae	MT621659.1	18S	AATCTATACGC
	217. https://doi.org/10.2903/j.efsa.2024.8970	Matsucoccus matsumurae	MW541961.1	18S	CAAATCTATGC
	218. https://doi.org/10.1007/BF02371987	Neomargarodes chondrillae	MH051908.1	18S	TTTGAAAGACC
	219. https://doi.org/10.7550/rmb.31286	Llaveia axin	HQ893805.1	12S	TTTGATAATGC
	220. https://doi.org/10.3958/059.049.0433	Insignorthezia insignis	HQ893803.1	12S	TTATTTTTTGG
	221. https://doi.org/10.1016/j.jobaz.2012.02.001	Phoenicococcus marlatti	KT199024.1	18S	CAAGTCATTGC
	222. https://doi.org/10.3897/BDJ.2.e1042	Coccidohystrix insolita	MW541948.1	18S	TCCGTATTGCC
	223. https://doi.org/10.2903/j.efsa.2021.6928	Crisicoccus pini	HG810185.1	16S	AGATTGTTACC
	224. https://doi.org/10.2903/j.efsa.2024.8740	Crisicoccus matsumotoi	KP692254.1	28S	TCCGTTTCCCC
	225. https://doi.org/10.1021/acs.jafc.4c05469	Delottococcus aberiae	KP771926.1	28S	CGTCATCTTCC
	226. https://doi.org/10.13140/RG.2.2.21810.81609	Delottococcus confusus	KP771927.1	28S	TACACGTCTCC
	227. https://doi.org/10.1590/S0085-56262009000100029	Dysmicoccus texensis	KY565032.1	28S	GTCATCTTCCC
	228. https://doi.org/10.1093/jee/toy344	Dysmicoccus lepelleyi	KX015061.1	28S	CTTCGCTTACC
	229. https://doi.org/10.46505/IJBI.2021.3112	Ferrisia virgata	MZ264180.1	28S	AACTCTTCAGC
	230. https://doi.org/10.1673/2006_06_23.1	Ferrisia malvastra	AY427383.1	28S	AAGATAAAAGC
	231. https://doi.org/10.1038/s42003-022-03956-y	Formicococcus polysperes	MZ920113.1	ITS2	TTTTTTTCGGC
	232. https://doi.org/10.1093/jisesa/iew043	Hypogeococcus pungens	AY427426.1	28S	CTTTCTTTTCC
	233. https://doi.org/10.1007/s12600-012-0260-2	Nipaecoccus nipae	AY427421.1	28S	TCCATTCATGC
	234. https://doi.org/10.1021/acs.jafc.0c07543	Nipaecoccus viridis	AY427393.1	28S	TTTTCCAAGCC
	235. https://doi.org/10.1007/s10584-017-1917-0	Oracella acuta	JF965418.1	28S	TTTCGCAGTCC
	236. https://doi.org/10.4001/003.018.0207	Paracoccus burnerae	AY426076.1	18S	TCCATTGATCC
	237. https://doi.org/10.2903/j.efsa.2023.7899	Paracoccus marginatus	AY427410.1	28S	TCTTTCCCCGC
	238. https://doi.org/10.1371/journal.pone.0047675	Phenacoccus manihoti	KY271371.1	28S	CTCTGAATTCC
	239. https://doi.org/10.1111/epp.12109	Phenacoccus madeirensis	AY427389.1	28S	TCTCGTAATCC
	240. https://doi.org/10.24154/jhs.v7i1.405	Phenacoccus parvus	MW542039.1	28S	CCTTTAGTACC
	241. https://doi.org/10.13140/2.1.4362.0165	Phenacoccus peruvianus	JF714177.1	28S	ATTAACGTACC
	242. https://doi.org/10.1303/aez.2006.573	Phenacoccus solani	KJ620514.1	28S	CTCGTTAATCC
	243. https://doi.org/10.1007/s13355-016-0452-1	Planococcus kraunhiae	MW542110.1	28S	AAGTAAATAGG
	244. https://doi.org/10.3389/fpls.2022.1016737	Planococcus lilacinus	KP692405.1	28S	TTACACGTACC
	245. https://doi.org/10.1603/ec12097	Planococcus minor	JF965414.1	28S	CTTCTTTCCCC
	246. https://doi.org/10.2298/PIF1401067G	Planococcus vovae	OR536427.1	28S	TAGTTTCTCGG
PSYLLOIDEA
	247. https://doi.org/10.1080/00222939100770791	Aphalaroida inermis	MG988580.1	18S	CACAGTTATCC
	248. https://doi.org/10.1080/00779962.2007.9722151.	Arytainilla spartiophila	MG988581.1	18S	AACCCTAATCC
	249. https://doi.org/10.11646/zootaxa.3646.2.2	Cornopsylla rotundiconis	MH758086.1	28S	TGCCCTTTTGC
	250. https://doi.org/10.1111/jen.12937	Diaphorina communis	MH042733.1	16S	GCTGTTATCCC
	251. https://doi.org/10.1111/j.1440-6055.1976.tb01714.x	Acizzia dodonaeae	MG195297.1	18S	CTAATAAAAGC
	252. https://doi.org/10.1371/journal.pone.0214220	Acizzia errabunda	MG195288.1	18S	CTAGAATTACC
	253. https://doi.org/10.13140/RG.2.2.22942.72008	Acizzia sasakii	MK039563.1	16S	ATTTCTCTTCC
	254. https://doi.org/10.11646/zootaxa.4362.1.4	Cacopsylla bidens	MK039580.1	16S	TCATACAAGCC
	255. https://doi.org/10.3897/BDJ.10.e85094	Cacopsylla burckhardti	MK039571.1	16S	ATAAAACACGC
	256. https://doi.org/10.11646/zootaxa.4362.1.4	Cacopsylla chinensis	LC513963.1	16S	TCTTATCGTCC
	257. https://doi.org/10.1080/23802359.2021.1875908	Cacopsylla citrisuga	MH053225.1	16S	ACTATCACCCC
	258. https://doi.org/10.1186/s12866-020-01895-4	Cacopsylla heterogena	MH053234.1	16S	AAAAATTATGC
	259. https://doi.org/10.3390/agronomy13112710	Cacopsylla jukyungi	LC513968.1	16S	AATTCTATAGG
	260. https://doi.org/10.5281/zenodo.14578	Cacopsylla mali	AF367822.1	12S	GATAAAATACC
	261. https://doi.org/10.5281/zenodo.14578	Cacopsylla peregrina	MT038953.1	28S	TGCTTAAATCC
	262. https://doi.org/10.34101/actaagrar/74/1660	Cacopsylla pruni	DQ778635.1	18S	GTACTCATTCC
	263. https://doi.org/10.5281/zenodo.5806004	Cacopsylla pulchra	MT038955.1	28S	TATTAATATGC
	264. https://doi.org/10.3390/agronomy14040668.	Cacopsylla pyri	MK039584.1	16S	AAATTATAAGG
	265. https://doi.org/10.11646/zootaxa.4362.1.4	Cacopsylla pyricola	MK039589.1	16S	TCTGTTCAACC
	266. https://doi.org/10.11646/zootaxa.4362.1.4	Cacopsylla pyrisuga	AB721006.1	16S	ACATTTTCCCC
	267. https://doi.org/10.5281/zenodo.14578	Cacopsylla ulmi	MT038960.1	28S	TTAAATCCACC
	268. https://doi.org/37. 10.5281/zenodo.5806004.	Livilla horvathi	AF367826.1	12S	CTTTTAAATCC
	269. http://dx.doi.org/10.5252/z2015n1a13	Livilla pyrenaea	AF367832.1	12S	TTAATAATTCC
	270. https://doi.org/10.4289/0013-8797.119.1.162	Livilla variegata	AF367837.1	12S	ATCCTATTTCC
	271. https://doi.org/10.13140/RG.2.2.14711.39848	Psylla alni	MG988605.1	18S	ACTTTGCTTGC
	272. https://doi.org/10.13140/RG.2.2.14711.39848	Psylla alniformosanaesuga	MH758107.1	28S	GGTATTTCACC
	273. https://doi.org/10.1111/epp.12636	Psylla buxi	MG988606.1	18S	CTTTTACTTCC
	274. https://doi.org/37. 10.5281/zenodo.5806004.	Psylla foersteri	MG988583.1	18S	AATACGAATGC
	275. https://doi.org/10.3897/travaux.66.e98619	Spanioneura fonscolombii	MG988609.1	18S	ATCAAGTTTGG
	276. https://doi.org/10.21203/rs.3.rs-1641424/v1	Bactericera cockerelli	MW804913.1	18S	AATTTGCGTGC
	277. https://doi.org/10.2903/j.efsa.2021.6357	Diaphorina citri	MG997020.1	16S	TTGGTATGTCC
	278. https://doi.org/10.3389/fpls.2022.1089762	Trioza erytreae	MT416551.1	12S	GTAAATTTACC
	279. https://doi.org/10.3897/zookeys.319.4316	Livia junci	NC_038137.1	16S	TAATTTTATGC
	280. https://doi.org/10.3390/insects15010019	Bactericera gobica	MH757801.1	18S	TACGCTATTGG
	281. https://doi.org/10.3844/ajabssp.2008.686.688	Bactericera tremblayi	MT038925.1	5.8S	TTCGACTTGCC
	282. https://doi.org/10.1111/eea.12565	Bactericera trigonica	MT038942.1	ITS2	TTTGGAATAGC
	283. https://doi.org/10.1371/journal.pone.0257031	Trioza eugeniae	U06482.1	18S	AAATTGCAAGC
	284. https://doi.org/10.1371/journal.pone.0257031	Trioza adventicia	MG195401.1	18S	CTTGTAAAAGG
HETEROPTERA
	285. https://doi.org/10.1093/ismejo/wrae097	Nezara viridula	AB020419.1	16S	GATTTAAGTGC
	286. https://doi.org/10.1371/journal.pone.0108746	Oncopeltus fasciatus	LN623664.1	28S	GTTTTCCACCC
	287. https://doi.org/10.3390/insects14110843	Pyrrhocoris apterus	KX821803.1	12S	GTAAAATTACC
	288. https://doi.org/10.3897/BDJ.7.e36453	Acanthocoris sordidus	AB240575.1	16S	TAATACCATGG
	289. https://doi.org/10.1073/pnas.1202111109	Lygaeus kalmii	KC425216.1	18S	GTTCTCATTCC
	290. https://doi.org/10.3390/insects10060167	Apolygus lucorum	KU049703.1	16S	TAAATCATAGG
	291. https://doi.org/10.1371/journal.pone.0059000	Adelphocoris lineolatus	KJ461200.1	18S	GTTTACTGAGC
	292. https://doi.org/10.4269/ajtmh.2012.11-0237	Triatoma infestans	MW893248.1	16S	TAAACAAAAGG
	293. https://doi.org/10.3390/insects13060511	Riptortus pedestris	GU145271.1	16S	CCTTTCATACC
	294. https://doi.org/10.1264/jsme2.ME19011	Coreus marginatus	MW845175.1	12S	AATCTCTATGC
	295. https://doi.org/10.3390/insects12090807	Lygus lineolaris	KP230905.1	28S	TGAATCTCCCC
	296. https://doi.org/10.1673/031.011.13801	Eurygaster integriceps	KP890857.1	18S	CCAATAGATCC
	297. https://doi.org/10.1673/031.013.5001	Aelia acuminata	MT265584.1	18S	TTTAAGTCAGC
	298. https://doi.org/10.1093/jisesa/ieae025	Dolycoris baccarum	JQ029133.1	12S	TAATAATAGGG
	299. https://doi.org/10.1038/s41598-023-37705-w	Eurydema ventralis	MT265521.1	28S	GTTTTACACCC
	300. https://doi.org/10.1038/s41598-023-37705-w	Eurydema oleracea	NC_044764.1	12S	TAATTTCTAGG
	301. https://doi.org/10.2903/j.efsa.2022.7091	Bagrada hilaris	OP168132.1	16S	AATGATTATGC
	302. https://doi.org/10.1371/journal.pone.0152205	Corythucha ciliata	MZ241573.1	18S	GATTGTGTTGC
	303. https://doi.org/10.3390/insects11020129	Stephanitis pyri	KF661826.1	28S	TTGTAACACCC
	304. https://doi.org/10.1264/jsme2.ME22042	Leptoglossus occidentalis	ON331774.1	18S	GCTTTTTAACC
	305. https://doi.org/10.3390/insects10100333	Leptoglossus zonatus	KF057195.1	28S	GTTGTATTAGC
	306. https://doi.org/10.1093/jisesa/iey021	Anasa tristis	MW883642.1	16S	AAATCTAAAGG
	307. https://doi.org/10.1038/s41598-022-16295-z	Nezara antennata	MT265637.1	18S	AAACTATGACC
	308. https://doi.org/10.3389/fpls.2022.804839	Piezodorus guildinii	JX425419.1	16S	TTCATACAAGC
	309. https://doi.org/10.3897/BDJ.7.e36453	Taylorilygus apicalis	MH432249.1	28S	TTTTGACACCC
	310. https://doi.org/10.1002/ece3.510	Creontiades dilutus	NC_030257.1	12S	AGATTTATTGG
	311. https://doi.org/10.1080/09670874.2017.1403059	Graphosoma lineatum	U88339.1	18S	TAACTACTGGC
	312. https://doi.org/10.1111/jen.13206	Palomena prasina	JQ029154.1	16S	CTCATTCAACC
	313. https://doi.org/10.55446/IJE.2022.805	Eurydema dominulus	KC155935.1	16S	CTCTCTCATCC
	314. https://doi.org/10.1038/s41598-023-37705-w	Eurydema ornata	MT265620.1	18S	TGCCTGTTACC
	315. https://doi.org/10.15835/hpm.v31i1-2.14746	Oxycarenus lavaterae	LN623676.1	28S	CTCTCTGATCC
	316. https://doi.org/10.5073/JABFQ.2015.088.003	Eurygaster maura	JQ029150.1	16S	CGCTGTTATCC
	317. https://doi.org/10.17660/ActaHortic.2014.1052.38	Gonocerus acuteangulatus	MW845231.1	12S	TATATAACACC
	318. https://doi.org/10.1155/2012/435640	Trigonotylus caelestialium	MZ241805.1	28S	CTGTCTTAAGC
	319. https://doi.org/10.1080/09583157.2020.1828279	Dicyphus errans	KY274682.1	16S	AAAATTATAGG
	320. https://doi.org/10.1080/23802359.2019.1674732	Cletus punctiger	AY627323.1	18S	TCAATAGTACC
	321. https://doi.org/10.21203/rs.3.rs-4239869/v1	Apolygus spinolae	GU194615.1	18S	TCATCGATACC
	322. https://doi.org/10.1088/1755-1315/913/1/012012	Riptortus linearis	AY665547.1	18S	ACATACTTGGC
	323. https://doi.org/10.1002/ps.5518	Pseudatomoscelis seriatus	OR229249.1	28S	TTCAGCTTTGC
	324. https://doi.org/10.3897/neobiota.90.113421	Halymorpha halys	KF273403.1	28S	ATCTCTGATCC
	325. https://doi.org/10.18474/0749-8004-44.1.1	Acrosternum hilare	MK648451.1	16S	TAAGCTTCTGC
	326. https://doi.org/10.1002/ps.6821	Dichelops furcatus	KU853750.1	28S	CACTTTTAAGC
	327. https://doi.org/10.1007/s10886-012-0144-4	Edessa meditabunda	KF035964.1	16S	TCTATAAATCC
	328. https://doi.org/10.18016/ksutarimdoga.vi.1388051	Aelia rostrata	MT265398.1	16S	AATTTTAAAGG
	329. https://doi.org/10.1007/s10340-021-01445-5	Oebalus ypsilongriseus	KC537028.1	16S	AATTTTAATCC
	330. https://doi.org/10.1590/S1519-566X2007000600022	Arvelius albopunctatus	MT073065.1	18S	TTTAATTCAGC
	331. https://doi.org/10.1002/ps.8854	Brachyplatys subaeneus	KY495723.1	28S	TTAAATTCGGC
	332. https://doi.org/10.1303/jjaez.2003.159	Plautia crossota	HQ630645.1	18S	CTGCTTTAAGC
	333. https://doi.org/10.1303/jjaez.2021.13	Plautia stali	LC007117.1	16S	ATTAAATTACC
	334. https://doi.org/10.1093/jee/toad177	Leptoglossus clypealis	MW845137.1	12S	ACCAATCAACC
	335. https://doi.org/10.4039/Ent128777-4	Leptoglossus corculus	MW838831.1	28S	TCCCTTTCGCC
	336. https://doi.org/10.1093/aesa/83.4.753	Leptoglossus fulvicornis	MW838585.1	18S	ATTTAAGAGCC
	337. https://doi.org/10.3958/0147-1724-33.2.161	Leptoglossus phyllopus	MW838836.1	28S	ACAACTCTCCC
	338. https://doi.org/10.1603/AN13084	Narnia femorata	MG490147.1	18S	GATATGTCACC
	339. https://doi.org/10.1590/S0103-84782001000300003	Phthiacnemia picta	KC537014.1	16S	CTATCTAAAGG
	340. https://doi.org/10.32473/EDIS-IN582-2003	Spartocera batatas	MW838671.1	18S	ATTTGATAGCC
	341. https://doi.org/10.5281/zenodo.8368595	Arocatus longiceps	LN623643.1	28S	CTCAAACTCCC
	342. https://doi.org/10.21661/r-471192	Arocatus melanocephalus	KJ461273.1	18S	TAGTTAATAGG
	343. https://doi.org/10.21661/r-117698	Kleidocerys resedae	LN623655.1	28S	ATCCTCCCTCC
	344. https://doi.org/10.1038/srep11042	Adelphocoris suturalis	GU194529.1	16S	AAATTTAAAGG
	345. https://doi.org/10.1007/s10340-009-0277-6	Closterotomus trivialis	MF667144.1	28S	GTTTTGACACC
	346. https://doi.org/10.1007/s10340-016-0765-4	Campylomma verbasci	KF273138.1	28S	ATATGGACCC
	347. https://doi.org/10.3923/je.2007.324.330	Lygus elisus	AF473541.1	16S	AGAAAGTTTGC
	348. https://doi.org/10.1002/ps.6303	Lygus rugulipennis	GU194565.1	16S	TCTTATTATCC
	349. https://doi.org/10.1093/jee/toz340	Euschistus heros	KY440164.1	18S	TTGTCACTACC
	350. https://doi.org/10.1007/s13744-019-00706-4	Thyanta perditor	MT265670.1	18S	CATATATCCGG
	351. https://doi.org/10.1590/s1519-69842006000300010	Oebalus poecilus	KC796346.1	28S	CTTCATTGCGC
	352. https://doi.org/10.1016/j.jinsphys.2020.104038	Lygus hesperus	KP231017.1	28S	CGAATAAATCC
	353. https://doi.org/10.3390/insects15100762	Lygus pratensis	NC_037926.1	12S	AGATCTATTGG
	354. https://doi.org/10.1038/s41598-025-95657-9	Corythucha marmorata	KF661732.1	16S	TTATCATAGGG
	355. https://doi.org/10.1603/en11323	Stephanitis takeyai	KF661808.1	28S	TTTGTAACACC
	356. https://doi.org/10.1079/cabicompendium.30340	Leptoglossus gonagra	MW845149.1	12S	TTAATTCACCC
	357. https://doi.org/10.1017/s1431927619015010	Scaptocoris castanea	MG253280.1	16S	CTAAATTGACC
	358. https://doi.org/10.3390/insects14030237	Megacopta cribraria	AB240165.1	16S	ATAATATAAGC
	359. https://doi.org/10.1017/s0007485318000950	Creontiades pacificus	KC852033.1	28S	CATTTGATACC
	360. https://doi.org/10.1603/en11032	Stenotus rubrovittatus	GU194680.1	18S	AGATTTCAGCC
	361. https://doi.org/10.1002/ps.6797	Teleonemia scrupulosa	KF661827.1	28S	TTTGTAACGCC
AUCHENORRHYNCHA
	362. https://doi.org/10.2903/j.efsa.2017.5037	Hishimonus phycitis	MT981712.1	16S	TAATTATGTCC
	363. https://doi.org/10.3389/fmicb.2023.1162113	Nilaparvata lugens	AF158034.1	16S	AATATAATACC
	364. https://doi.org/10.1002/arch.21992	Laodelphax striatellus	PQ500564.1	18S	CTTAATGTGGC
	365. https://doi.org/10.7717/peerj.9320	Sogatella furcifera	HM017358.1	28S	CTACTACCACC
	366. https://doi.org/10.3390/ijms221910696	Nephotettix cincticeps	AY830909.1	16S	TATCTTAATCC
	367. https://doi.org/10.1088/1755-1315/807/2/022107	Nephotettix virescens	KR230142.1	28S	TTTTCTATCGG
	368. https://doi.org/10.3390/v16081305	Empoasca fabae	AF304640.1	28S	ATTAGATTAGG
	369. https://doi.org/10.1002/ps.8659	Amrasca biguttula	PP853663.1	12S	AATAAAAGAGC
	370. https://doi.org/10.1127/entomologia/2020/0977	Scaphoideus titanus	MT345578.1	16S	ATATGAACTCC
	371. https://doi.org/10.1603/en10115	Homalodisca vitripennis	AY869803.1	16S	GTAAATTCTGC
	372. https://doi.org/10.1128/AEM.01464-15	Macrosteles quadrilineatus	AF051288.1	12S	TACTATTCAGG
	373. https://doi.org/10.1094/pdis-10-15-1189-re	Circulifer tenellus	LC670614.1	28S	AATTTCATTGC
	374. https://doi.org/10.1002/ps.6842	Dalbulus maidis	AF051283.1	12S	TAACTAATACC
	375. https://doi.org/10.4137/IJIS.S13029	Cofana spectra	GQ122082.1	16S	GAAAATCTACC
	376. https://doi.org/10.3791/62417	Peregrinus maidis	HM017225.1	18S	ACTAAGATCCC
	377. https://doi.org/10.1046/j.1440-6055.2002.00268.x	Idioscopus clypealis	KX268303.1	28S	TTACCACCTGC
	378. https://doi.org/10.3390/insects14090734	Maiestas dorsalis	MT988057.1	28S	AGTTTACCACC
	379. https://doi.org/10.3390/insects11020080	Spissistilus festinus	AF183246.1	28S	TCCATCTAAGG
	380. https://doi.org/10.46429/jaupr.v57i4.10729	Umbonia crassicornis	AF187463.1	28S	TCTGTGAAACC
	381. https://doi.org/10.1590/0103-8478cr20170152	Quesada gigas	MG953135.1	18S	TTATGACTCGC
	382. https://doi.org/10.1128/mra.00446-22	Exitianus exitiosus	L13018.1	16S	AAGAATTCTGC
	383. https://doi.org/10.1093/jee/tov063	Empoasca vitis	JQ886745.1	16S	TAATCATTGGG
	384. https://doi.org/10.1111/epp.12610	Philaenus spumarius	AY744779.1	18S	CATCTTTCCGG
	385. https://doi.org/10.3390/insects12040340	Poophilus costalis	GQ405992.1	18S	CTTAATTTGGC
	386. https://doi.org/10.1007/s00203-003-0630-8	Dalbulus elimatus	KU722545.1	16S	TGTCTCAGTCC
	387. https://doi.org/10.1002/jmor.1051720310	Macrosteles fascifrons	L12998.1	16S	TATCATTGGGC
	388. https://doi.org/10.3390/insects15040296	Empoasca flavescens	AY830906.1	28S	CACTATCAAGC
	389. https://doi.org/10.1016/j.phytochem.2021.112848	Neophilaenus campestris	KP410336.1	ITS2	TTTACCGAACC
	390. https://doi.org/10.1080/23802359.2021.1911705	Stictocephala bisonia	PV207583.1	12S	CCTATTTTAGG
	391. https://doi.org/10.1094/pdis.2003.87.10.1255	Homalodisca coagulata	AY875213.1	28S	TGAAATAATGC
	392. https://doi.org/10.3390/insects11060345	Metcalfa pruinosa	KX702853.1	18S	TTGTACGATCC
	393. https://doi.org/10.2903/j.efsa.2020.6224	Haplaxius crudus	HM017369.1	28S	ATTTTCAGGGC
	394. https://doi.org/10.1038/s41598-020-70150-7	Ommatissus lybicus	KP822941.1	28S	TGAATCTCTCC
THYSANOPTERA
	395. https://doi.org/10.1017/S0007485319000762	Thrips tabaci	AB904212.1	ITS1	CACCTTTCCCC
	396. https://doi.org/10.1371/journal.pone.0199404	Thrips palmi	OR350842.1	ITS2	ATATATGATCC
	397. https://doi.org/10.1515/aeuc-2016-0005	Thrips simplex	KM877312.1	ITS2	TCTTAGTGACC
	398. https://doi.org/10.1111/j.1365-2338.2008.01201.x	Thrips hawaiiensis	AB904201.1	ITS1	GTTTTGTACGG
	399. https://doi.org/10.3390/insects12060502	Thrips flavus	AM932174.1	ITS2	TGTTAAACACC
	400. https://doi.org/10.1303/jjaez.2014.275	Thrips nigropilosus	AM932193.1	ITS2	TTGATCCAACC
	401. https://doi.org/10.18520/cs/v122/i2/211-213	Thrips parvispinus	OR141114.1	18S	TATCTGATCGC
	402. https://doi.org/10.1093/jipm/pmab045	Frankliniella fusca	AB972997.1	ITS2	AATCTCGTTCC
	403. https://doi.org/10.3390/insects13060493	Frankliniella insularis	GU980274.1	28S	TCTTCCCAAGC
	404. https://doi.org/10.3390/insects14070643	Frankliniella occidentalis	AJ308591.1	ITS2	GTTTCGTCTCC
	405. https://doi.org/10.1653/024.097.0206	Frankliniella schultzei	GQ343259.1	ITS1	TTTTTTAAGGC
	406. https://doi.org/10.32473/edis-in1236-2019	Frankliniella tritici	AB973010.1	ITS2	TTAAGAGTACC
	407. https://doi.org/10.1093/jisesa/iey028	Scirtothrips aurantii	DQ075048.1	ITS1	AGTTTTTACGG
	408. https://doi.org/10.1371/journal.pone.0123747	Scirtothrips dorsalis	DQ075059.1	ITS1	CTGAATTCCCC
	409. https://doi.org/10.1079/BER2002169	Scirtothrips perseae	DQ075029.1	ITS2	CAATCTCGTCC
	410. https://doi.org/10.1111/afe.12233	Anaphothrips obscurus	AB973073.1	ITS2	AATTATATTGG
	411. https://doi.org/10.1038/s41598-025-98466-2	Fulmekiola serrata	KC505476.1	16S	TTTCGATTTGG
	412. https://doi.org/10.33338/ef.84532	Limothrips denticornis	KT204371.1	18S	ATTACCTCTGG
	413. https://doi.org/10.1038/s41597-023-02164-5	Megalurothrips usitatus	AB904185.1	ITS1	TCGGACTTTCC
	414. https://doi.org/10.1603/en13002	Pezothrips kellyanus	KC513094.1	28S	TTTCAATGTCC
	415. https://doi.org/10.14719/pst.5230	Pseudodendrothrips mori	KC513000.1	18S	GTCATCTTTCC
	416. https://doi.org/10.32473/edis-IN1364-2022	Caliothrips fasciatus	JQ609601.1	28S	AACAATAAAGC
	417. https://doi.org/10.25100/socolen.v50i1.13135	Caliothrips phaseoli	AB972981.1	28S	GTTTCTTTTCC
	418. https://doi.org/10.1093/ee/nvac021	Heliothrips haemorrhoidalis	KC512969.1	18S	GTCAACTTCCC
	419. https://doi.org/10.1007/s41348-023-00751-7	Hercinothrips femoralis	AJ308596.1	28S	ATTGTTTTTGG
	420. https://doi.org/10.1079/cabicompendium.62325634	Thrips setosus	AB277570.1	28S	TTTCGTAAGCC
	421. https://doi.org/10.2903/j.efsa.2018.5189	Scirtothrips citri	KX610660.1	28S	CATGAATCTCC
	422. https://doi.org/10.3390/insects13010061	Mycterothrips glycines	HM246165.1	16S	CTTTCTTTAGG
	423. https://doi.org/10.1080/23802359.2023.2169572	Stenchaetothrips biformis	KC513010.1	18S	CTTGCACTTGC
	424. https://doi.org/10.1002/arch.22086	Aptinothrips stylifer	KJ436475.1	28S	CTATACTCAGC
	425. https://doi.org/10.3390/insects13010061	Frankliniella tenuicornis	KM886245.1	ITS1	TGCTTGTTAGC
	426. https://doi.org/10.1093/jee/toy237	Frankliniella williamsi	AB904183.1	ITS1	TTTGCTTTCGC
	427. https://doi.org/10.3390/insects15090699	Thrips fuscipennis	AB904198.1	ITS1	TTTTGTACGGG
	428. https://doi.org/10.30843/nzpp.2008.61.6856	Thrips major	KM877309.1	ITS2	TACGGTATACC
	429. https://doi.org/10.3897/zookeys.330.5939	Thrips physapus	AB973055.1	ITS2	TCACAGTTTGG
	430. https://doi.org/10.3897/zookeys.504.9576	Thrips trehernei	KM877313.1	ITS2	ATTGCAATCGC
	431. https://doi.org/10.3897/zookeys.504.9576	Thrips vulgatissimus	AB973064.1	ITS2	CTTGTCATCGC
	432. https://doi.org/10.3897/zookeys.549.6889	Selenothrips rubrocinctus	KC513107.1	28S	CAATGACTTGC
	433. https://doi.org/10.2903/j.efsa.2022.7337	Microcephalothrips abdominalis	AB904191.1	18S	GGTCATCTTCC
	434. https://doi.org/10.1673/031.006.4501	Frankliniella cephalica	AB904174.1	18S	CATTCAATCGG
	435. https://doi.org/10.1093/jisesa/ieu167	Frankliniella intonsa	LC662016.1	ITS2	TCTCTATACGG
	436. https://doi.org/10.3390/toxins14070502	Limothrips cerealium	AJ308593.1	ITS2	TCTTAAAACGC
	437. https://doi.org/10.1371/journal.pone.0276865	Frankliniella gardeniae	KY605027.1	28S	CCGTTTACACC
	438. https://doi.org/10.3390/insects13010061	Parthenothrips dracaenae	AJ308595.1	ITS2	TTTCAAATGGG
	439. https://doi.org/10.1093/jisesa/ieu135	Gynaikothrips uzeli	MK940484.1	16S	TATTTTAAACC
	440. https://doi.org/10.3390/insects11120887	Gynaikothrips ficorum	KC513064.1	28S	ATTAATATGCC
	441. https://doi.org/10.3390/insects15030211	Echinothrips americanus	AB972994.1	5.8S	TTCATCGACCC
	442. https://doi.org/10.3390/insects15090699	Thrips australis	KC513117.1	28S	CTACATTTCCC
	443. https://doi.org/10.1038/s41598-024-63985-x	Taeniothrips inconsequens	KC513112.1	28S	TTTTCCTCCCC
	444. https://doi.org/10.3390/insects13010061	Dendrothrips ornatus	PX262664.1	16S	AATCCTATCGG
	445. https://doi.org/10.3390/insects13010061	Rhipiphorothrips cruentatus	MN072396.1	16S	TAGTTAAACCC
	446. https://doi.org/10.1093/jisesa/ieae086	Dictyothrips betae	EU704684.1	28S	TTAGGTTTCGC
	447. https://doi.org/10.3390/insects13010061	Sericothrips staphylinus	KC513108.1	28S	CTTTTCCTCCC
	448. https://doi.org/10.3390/insects13010061	Aptinothrips rufus	AB972972.1	5.8S	CTTCATCGACC
	449. https://doi.org/10.3390/insects13010061	Anaphothrips sudanensis	AB973066.1	ITS2	ATTTATATTGG
	450. https://doi.org/10.1016/j.compag.2022.107462	Thrips imaginis	AB775432.1	5.8S	CATTTCGCTGC
	451. https://doi.org/10.3390/insects13080670	Thrips brevicornis	NC_087731.1	16S	CAAATGTTTGG
	452. https://doi.org/10.33936/latecnica.v13i2.4640	Chaetanaphothrips signipennis	MK056207.1	18S	AGTTTGACACC
	453. https://doi.org/10.1093/jee/toaa311	Frankliniella bispinosa	GU980272.1	28S	GTAATCTCACC
TETRANYCHIDAE
	454. https://doi.org/10.1016/j.ijbiomac.2024.138982	Amphitetranychus viennensis	AB926293.1	18S	AAGGAATATCC
	455. https://doi.org/10.11158/saa.3.1.8	Aponychus corpuzae	AB926233.1	18S	ATGTAAATTGG
	456. https://doi.org/10.1093/jee/toz294	Bryobia praetiosa	AB926228.1	18S	GGTTTTTGTGC
	457. https://doi.org/10.13140/RG.2.1.3259.6967	Bryobia rubrioculus	MW530361.1	ITS2	TGATACTCCCC
	458. https://doi.org/10.1023/A:1021116121604	Eotetranychus asiaticus	AB926259.1	18S	ATGGAATATCC
	459. https://doi.org/10.1603/EC12304	Eotetranychus lewisi	JF774172.1	ITS2	TGTTTCTCAGG
	460. https://doi.org/10.1023/A:1021185112192	Eotetranychus pruni	AB926356.1	28S	TATGCTACACC
	461. https://doi.org/10.4067/S0718-58392021000200228	Eutetranychus africanus	AB926232.1	18S	AATTACAATGC
	462. https://doi.org/10.24349/pv5i-1hqj	Eutetranychus banksi	KP642050.1	5.8S	ATTATCGTAGC
	463. https://doi.org/10.1007/s10493-011-9476-y	Eutetranychus orientalis	DQ656470.1	ITS2	TATACTAAACC
	464. http://dx.doi.org/10.22073/pja.v8i2.43946	Eutetranychus palmatus	DQ656454.1	ITS2	TATACTGAACC
	465. https://doi.org/10.1093/ee/24.4.841	Mononychellus progresivus	X79902.1	ITS2	TGTATAGTTGG
	466. https://doi.org/10.1673/031.011.13601	Oligonychus afrasiaticus	OQ185412.1	ITS	AGTATACAAGC
	467. https://doi.org/10.3390/insects16010096	Oligonychus biharensis	AB926277.1	18S	ATGTAAAAAGG
	468. https://doi.org/10.1007/s10493-014-9800-4	Oligonychus coffeae	AY750699.1	28S	ATCTAGACTCC
	469. https://doi.org/10.1093/jee/toae086	Oligonychus ilicis	AB926284.1	18S	ATGTAAAATGG
	470. https://doi.org/10.2903/j.efsa.2021.6927	Oligonychus mangiferus	DQ656486.1	ITS2	ATACATTTACC
	471. https://doi.org/10.2903/j.efsa.2017.5075	Oligonychus perditus	AB926288.1	18S	ATTACAATGCC
	472. https://doi.org/10.1007/s10493-016-0018-5	Oligonychus perseae	KY474116.1	28S	GTTTCACGTCC
	473. https://doi.org/10.1093/jee/toab029	Oligonychus punicae	KC352232.1	18S	AAAAAATAAGG
	474. https://doi.org/10.1093/jee/96.6.1675	Oligonychus ununguis	AB683728.1	28S	TTTTGCCAAGC
	475. https://doi.org/10.1111/1744-7917.13265	Panonychus citri	XR_008326918.1	28S	CTTAAATTCGG
	476. https://doi.org/10.1007/s12600-025-01291-6	Panonychus ulmi	AY750698.1	28S	TACTAGACTCC
	477. https://doi.org/10.1080/01647954.2010.519720	Petrobia harti	AY750696.1	28S	TTTCAATGCGC
	478. https://doi.org/10.1093/jee/toz157	Petrobia latens	AB926229.1	18S	AACATTCTTGG
	479. https://doi.org/10.1007/s10493-019-00402-3	Schizotetranychus brevisetosus	AB926337.1	28S	CCCACTTATGC
	480. https://doi.org/10.20546/IJCMAS.2019.809.081	Schizotetranychus lespedezae	AB926340.1	28S	CCAGTTATCCC
	481. https://doi.org/10.11158/saa.26.2.2	Schizotetranychus schizopus	AB926342.1	28S	TTTCTGACACC
	482. https://doi.org/10.3897/BIORISK.4.58	Stigmaeopsis celarius	AB926253.1	18S	TAAGAATATCC
	483. https://doi.org/10.1007/s10493-008-9147-9	Stigmaeopsis miscanthi	AB926255.1	18S	TTACAATGCCC
	484. https://doi.org/10.1007/s10493-022-00718-7	Stigmaeopsis nanjingensis	LC158888.1	18S	TATGAATATCC
	485. https://doi.org/10.1016/j.aspen.2014.07.008	Tetranychus bambusae	AB926385.1	28S	AGATTGATCCC
	486. https://doi.org/10.1016/j.pestbp.2024.106112	Tetranychus cinnabarinus	GU550646.1	28S	AGTCAAACTCC
	487. https://doi.org/10.1007/s10493-012-9645-7	Tetranychus evansi	AB926295.1	18S	AAAGAATATCC
	488. https://doi.org/10.1603/022.038.0323	Tetranychus kanzawai	AY750691.1	28S	ACCATTTGTCC
	489. https://doi.org/10.1590/1519-6984.176665	Tetranychus ludeni	AB926300.1	18S	TTACCCACTCC
	490. https://doi.org/10.3390/plants14121765	Tetranychus macfarlanei	AB926301.1	18S	AAACAGTATGC
	491. https://doi.org/10.7717/peerj.14064	Tetranychus merganser	AB926393.1	28S	TAATCAAAAGG
	492. https://doi.org/10.1590/s1519-566x2010000300002	Tetranychus mexicanus	OK632064.1	ITS2	TATATATCTCC
	493. https://doi.org/10.3389/fpls.2018.01466	Tetranychus neocaledonicus	AB926304.1	18S	ACTCCTTTCGG
	494. https://doi.org/10.1093/jee/toab110	Tetranychus pacificus	X99879.1	ITS2	TGCATATATGG
	495. https://doi.org/10.24349/men5-cyrn	Tetranychus phaselus	AB926307.1	18S	AAATATCGAGC
	496. https://doi.org/10.1038/s41597-024-03189-0	Tetranychus piercei	AY750694.1	28S	AAACCGATTCC
	497. https://doi.org/10.1303/aez.2004.675	Tetranychus pueraricola	AB926309.1	18S	AAAATAGAACC
	498. https://doi.org/10.1016/j.pestbp.2025.106827	Tetranychus truncatus	AY750692.1	28S	AACCCTTCTCC
	499. https://doi.org/10.1007/s10493-025-01077-9	Tetranychus turkestani	AB926311.1	18S	AAAAACTTTGG
	500. https://doi.org/10.1038/nature10640	Tetranychus urticae	AY750693.1	28S	ATTTCTTTTGC

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