The origination of most free-living animal species is predictable. ‘Light’ order in the producer trophic levels below is the key. Absent from the abiotic environment (pre-biotic level) and prokaryotes, ‘light’ order consists of differences in species composition between developed species-packed communities in which extinction may be the precursor of speciation, particularly ecologically comparable tropical lowland ones of Africa, Australia, India, and South America. Based on but not itself of matter and non-burnable, nil waste heat content ‘compensates’ for burned-up food energy. Where the amounts on pre-apical levels form an inverted pyramid like waste heat’s, occupancy of consumer levels from primary to apical is predictable (variation and selection assumed). Terminal non-inversion predicts post-apical vacancy. Examined communities were from grasslands (large carnivores, large grazing mammals, grasses) and woodlands (raptorial birds, insect-eating birds, butterflies, woody plants). Comparisons with Darwin’s and Lyell’s non-predictive theories of change are made. Implication for Gladyshevian thermodynamics v. Prigoginean dynamics is discussed. Linnaeus’s classificational system turns out to be rich with new and unsuspected content.