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Diversity, Distribution, and Ethnobotanical Uses of Ornamental Plants in a Bangladeshi University Campus

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08 September 2026

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09 September 2026

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Abstract
Ornamental plants constitute an essential component of urban green infrastructure, contributing to biodiversity conservation, ecological resilience, and the aesthetic and cultural identity of institutional spaces. In Bangladesh, ethnobotanical knowledge embedded in the ornamental flora of university campuses remains largely undocumented. This study aimed to document the taxonomic diversity, spatial distribution patterns, and ethnobotanical uses of ornamental plants across the 360-acre campus of Daffodil International University (DIU), Bangladesh. Field surveys were conducted from March 2024 to February 2025 using systematic quadrat sampling (5 m × 5 m, n = 240 quadrats). Semi-structured interviews were conducted with 103 informants comprising campus staff, local inhabitants, and students. Diversity was quantified using the Shannon–Wiener index, Simpson's index, and Importance Value Index (IVI). Ethnobotanical significance was evaluated using Use Value (UV) and Informant Consensus Factor (ICF). A total of 68 ornamental plant species were recorded across 38 families. Aceraceae was the most species-rich family (n = 10), followed by Poaceae (n = 5) and Asparagaceae (n = 4). Life form analysis revealed that herbs (25.0%) and shrubs (23.53%) dominated the campus flora. Shannon–Wiener diversity (H' = 3.42) and Simpson's index (1–D = 0.94) indicated high species diversity. Ethnobotanical documentation revealed six major use categories: medicinal, aesthetic, cultural/ritual, environmental, economic, and nutritional. The DIU campus harbors a rich and culturally significant ornamental flora. Strategic conservation planning and ethnobotanically informed landscaping are recommended to enhance both biodiversity and traditional knowledge preservation.
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Introduction

Ornamental plants, cultivated primarily for their aesthetic, ecological, and cultural attributes, have gained increasing scientific attention as important components of urban biodiversity (Nawar et al., 2022; Saha et al., 2022). In rapidly urbanizing countries such as Bangladesh, university campuses serve as semi-natural refuges that support diverse plant assemblages while simultaneously fulfilling landscaping and ecological functions (Liu et al., 2021). These institutional green spaces not only enhance the visual environment but also serve as repositories of traditional ethnobotanical knowledge that connects communities to their natural heritage (Tama et al., 2021).
The ethnobotanical dimensions of campus flora in South Asia remain poorly characterized, despite the growing recognition that green spaces in academic institutions contribute meaningfully to urban ecosystem services including carbon sequestration, air quality improvement, thermal regulation, and psychological well-being (Nawar et al., 2022). Previous investigations of ornamental plant diversity in Bangladesh have been largely restricted to botanical gardens (Shomrat & Uddin, 2025) or rural home gardens (Roy et al., 2013), leaving university campuses underrepresented in the scientific literature.
Daffodil International University (DIU), located in Ashulia, Savar, Dhaka, encompasses approximately 360 acres of landscaped grounds characterized by a subtropical climate, rendering it suitable for a diverse range of ornamental taxa. The institution has invested substantially in campus greening initiatives, creating an ideal natural laboratory for evaluating ornamental plant diversity and associated ethnobotanical knowledge. A systematic botanical survey of DIU’s campus flora is therefore both timely and scientifically justified.
The specific objectives of this study were: (i) to document and classify the ornamental plant species present on the DIU campus according to taxonomic affiliation and life form; (ii) to quantify species diversity and Importance Value Index (IVI) of dominant tree and shrub species; (iii) to examine the spatial distribution of ornamental plants across different campus zones; and (iv) to record and analyze ethnobotanical uses of the documented species among campus inhabitants and associated communities, using standardized ethnobotanical indices.

Material and Methods

Study Area Description

The study was conducted at Daffodil International University (DIU) campus, located at Birulia, Ashulia, Savar, Dhaka Division, Bangladesh (23°53’N, 90°16’E). The campus covers approximately 360 acres and is divided into functional zones including academic buildings, residential halls, administrative blocks, recreational grounds, botanical trails, and water bodies. The climate is humid subtropical (Köppen: Cwa) with a mean annual temperature of 25.8 °C and annual precipitation of approximately 2,000 mm, concentrated in the monsoon season (June–September). Soils are predominantly alluvial loam with moderate organic matter content, conducive to a broad spectrum of ornamental taxa.

Sampling Design and Field Surveys

Systematic quadrat sampling was employed across the campus, with 240 quadrats of 5 m × 5 m each established along transects at 50 m intervals, stratified across seven ecological zones: central gardens, peripheral borders, roadside plantings, water margins, shaded understories, open lawns, and building surrounds. All vascular plant taxa cultivated for ornamental purposes were recorded within each quadrat. Data collected included species identity, frequency, and relative abundance.

Plant Identification and Voucher Collection

Plant specimens were collected, pressed, and identified using standard taxonomic keys and regional floras (M. S. Khan, 1972; Prain, 1963). Scientific nomenclature was verified against the World Checklist of Selected Plant Families (http://wcsp.science.kew.org/) and the Plant List (www.theplantlist.org). Voucher specimens were deposited in the DIU Herbarium with accession numbers prefixed DIU-001 through DIU-068.

Ethnobotanical Data Collection

Semi-structured interviews were conducted with 103 informants including groundskeeping staff (n = 28), local residents adjacent to the campus (n = 42), faculty members with botanical interest (n = 17), and students from life science faculty (n = 16). Ethnobotanical data were collected following the established guidelines (Cotton, 2002; Martin, 2004). Each informant was asked to identify plants they recognized and to describe any medicinal, cultural, economic, or nutritional uses. Prior informed consent was obtained from all participants and ethical approval was granted by the DIU Institutional Review Board (Ref: DIU-AGS-IRB-2024-07).

Data Analysis

Taxonomic diversity was assessed by tabulating families, genera, and species. Life forms were classified following Raunkiaer’s system as adapted for tropical floras. Species diversity was calculated using the Shannon–Wiener index (H’) and Simpson’s diversity index (1 – D). The Importance Value Index (IVI) for trees and shrubs was computed as the sum of relative density (RD), relative frequency (RF), and relative dominance (RDom), with a maximum value of 300. Ethnobotanical significance was assessed using Use Value (UV = ΣUi/n) and the Informant Consensus Factor (ICF = [Nur – Nt] / [Nur – 1]), where Nur is the number of use reports and Nt is the number of species used for a given category. All statistical analyses were performed in R version 4.3.2 (R Core Team, 2023).

Results and Discussion

Taxonomic Diversity and Family Composition

A total of 68 ornamental plant species were documented across the DIU campus, belonging to 38 families and 63 genera (Table 1). This richness compares favorably with analogous studies conducted in South Asian university settings: a researcher recorded 54 ornamental species at Rajshahi University (H. M. Mahbubur Rahman, 2015), while another researcher documented 61 species in a New Delhi campus survey . The predominance of Arecaceae (10 species, 14.71%) in the present study reflects Bangladesh’s tropical affinity and the traditional preference for palm species in institutional landscaping, consistent with findings from Chittagong University (Akter et al., 2022). Other well-represented families included Poaceae (5 species), Asparagaceae (4 species), Euphorbiaceae (3 species), and Cupressaceae (3 species).

Life Form Spectrum

The life form spectrum revealed that herbs constituted the most abundant category (n = 17; 25.0%), followed by shrubs (n = 16; 23.53%), trees (n = 12; 17.65%), palms (n = 10; 14.71%), grasses, ferns and climbers combined (n = 12; 5.88% each), and cycad (n = 1; 1.47%) (Figure 1). The dominance of herbs is consistent with deliberate campus landscaping strategies that favor low-maintenance boundary plantings and hedgerows (Liu et al., 2021). The presence of four climber species, including Jasminum sp., Passiflora coccinea, Philodendron hederaceum, and Thunbergia coccinea, reflects the utilization of vertical greenery elements in campus architecture, a trend increasingly documented in South Asian institutional campuses (Alam et al., 2025).

Species Diversity Indices

The Shannon–Wiener diversity index (H’) for the full campus flora was 3.42, and Simpson’s diversity index (1 – D) was 0.94, both indicating high species evenness and richness. These values exceed those reported by a researcher from Jahangirnagar University (H’ = 2.98; S. A. Khan et al., 2021) and approach values recorded in managed botanical gardens in Bangladesh (H’ = 3.61; Shomrat & Uddin, 2025), suggesting that deliberate ornamental planting at DIU has generated a comparably diverse plant assemblage. Palms contributed disproportionately to diversity through the inclusion of rare and introduced taxa, particularly Pinanga speciosa and Trachycarpus princeps, which are seldom recorded in other Bangladeshi campus surveys.

Importance Value Index and Dominant Species

The Importance Value Index analysis of trees and shrubs identified Polyalthia longifolia (IVI = 52.5) as the dominant ornamental tree on the DIU campus, consistent with its widespread deployment along avenues and building perimeters as a wind-resistant columnar tree in Bangladesh (Table 2). Ficus benjamina ranked second (IVI = 44.4), valued for its extensive canopy, followed by Borassus flabellifer (IVI = 39.5), a culturally iconic palm with deep ethnobotanical significance in rural Bangladesh (Figure 2). High Use Values were recorded for Polyalthia longifolia (UV = 2.8), Hibiscus rosa-sinensis (UV = 2.3), and Ficus benjamina (UV = 2.4), reflecting their dual roles in landscape and traditional use.

Ethnobotanical Uses

Across 103 informants, six major ethnobotanical use categories were identified: medicinal (28 species; ICF = 0.62; mean UV = 1.92), aesthetic/ornamental (68 species; ICF = 0.81; UV = 2.41), cultural/ritual (14 species; ICF = 0.55; UV = 1.38), environmental (32 species; ICF = 0.68; UV = 1.74), economic (19 species; ICF = 0.59; UV = 1.55), and nutritional (11 species; ICF = 0.51; UV = 1.22) (Table 3, Figure 3). The aesthetic/ornamental category predictably recorded the highest ICF and UV, given the campus context. However, the substantial reporting of medicinal uses (41.2% of species) underscores the deep ethnobotanical knowledge retained by campus groundskeeping staff and adjacent community members.
Catharanthus roseus was noteworthy for its simultaneous role as an ornamental species and a source of the anticancer alkaloid’s vinblastine and vincristine, with five informants spontaneously citing its therapeutic use for diabetes management, consistent with its well-documented pharmacological profile (van Der Heijden et al., 2004). Hibiscus rosa-sinensis, was cited by 71% of informants for medicinal properties including hair growth promotion, wound healing, and fever management. Borassus flabellifer was the most frequently cited species for nutritional and economic uses, reflecting its deep integration into Bengali rural culture (Millat-E-Mustafa et al., 2002).

Ecological Significance and Conservation Implications

The diversity of ornamental plants at DIU contributes substantially to campus ecosystem services. The 32 species categorized under environmental uses include Dypsis lutescens and Dracaena fragrans, both scientifically validated indoor air purifiers (Wolverton et al., 1989), and Polyalthia longifolia, valued for noise reduction along major roads. The campus water margins support Thalia geniculata and Zantedeschia aethiopica, which serve as habitat for aquatic invertebrates, thereby contributing indirectly to campus faunal diversity. Leyland cypress (Cupressocyparis leylandii) and Platycladus orientalis function as effective windbreaks, reducing soil erosion on the campus periphery.
The recorded taxonomic richness, while impressive for an institutional campus, is subject to the inherent limitations of deliberate ornamental planting: exotic species dominate, and native flora representation is modest. Only 31 of the 68 recorded species are indigenous or naturalized in Bangladesh, a proportion that compares unfavorably with recommendations for ecologically sensitive campus landscaping (Chowdhury et al., 2021). Future planting initiatives should priorities native taxa that support pollinator communities and restore connectivity with regional forest fragments.

Conclusions

This study provides the first systematic documentation of the ornamental plant diversity, distribution, and ethnobotanical uses of the Daffodil International University campus, Bangladesh. Sixty-eight ornamental species distributed across 38 families were recorded, with Arecaceae exhibiting the highest family-level richness. High Shannon–Wiener (H’ = 3.42) and Simpson’s (1 – D = 0.94) diversity indices confirm that the campus sustains a rich and well-distributed ornamental flora. Ethnobotanical analysis identified six functional use categories, with medicinal uses recorded for 41.2% of species, highlighting the culturally significant plant knowledge embedded in this institutional landscape.
These findings have practical implications for campus biodiversity management. It is recommended that future landscaping incorporates a higher proportion of native ornamental species to enhance ecological connectivity and pollinator support. The ethnobotanical knowledge documented here, particularly regarding medicinal species such as Catharanthus roseus and Hibiscus rosa-sinensis, should be preserved through campus botanical trails with interpretive signage. Future research should assess the invertebrate and avian communities supported by the campus flora, and evaluate temporal changes in plant diversity in response to campus expansion activities.

Author Contributions

R.H.R. conceived and designed the study, conducted field investigations, collected and analyzed the data, interpreted the results, prepared the figures and tables, and wrote the main manuscript text. F.T. contributed to data collection, proofreading, and critical revision of the manuscript. All authors reviewed and approved the final manuscript.

Funding

No funding was received for this study.

Data Availability Statement

The data used in this study are available from the corresponding author upon reasonable request.

Acknowledgments

The authors express sincere gratitude to the administration of Daffodil International University for facilitating unhindered access to the campus during field surveys. Appreciation is extended to all 103 informants for generously sharing their botanical knowledge. The authors also acknowledge the Department of Botany herbarium staff for assistance with specimen curation. No external funding was received for this research.

Conflict of Interest

The authors declare that they have no conflict of interest.

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Figure 1. Life form spectrum of ornamental plant species on the DIU campus (n = 68).
Figure 1. Life form spectrum of ornamental plant species on the DIU campus (n = 68).
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Figure 2. IVI components (relative density, frequency, dominance) and Use Values (UV; red diamonds) for the top 10 ornamental tree and shrub species on the DIU campus. Maximum IVI = 300.
Figure 2. IVI components (relative density, frequency, dominance) and Use Values (UV; red diamonds) for the top 10 ornamental tree and shrub species on the DIU campus. Maximum IVI = 300.
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Figure 3. Ethnobotanical use category distribution on the DIU campus. (A) Number and percentage of species per use category. (B) Mean Use Value (UV) radar chart per category. n = 103 informants; 68 species.
Figure 3. Ethnobotanical use category distribution on the DIU campus. (A) Number and percentage of species per use category. (B) Mean Use Value (UV) radar chart per category. n = 103 informants; 68 species.
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Table 1. Ornamental plant species documented on the DIU campus, Bangladesh, with taxonomic classification, voucher accession numbers, and ethnobotanical uses (n = 68 species).
Table 1. Ornamental plant species documented on the DIU campus, Bangladesh, with taxonomic classification, voucher accession numbers, and ethnobotanical uses (n = 68 species).
SL Common Name Scientific Name Family Life Form Voucher No. Ethnobotanical Uses
1 Chinese togor Tabernaemontana divaricata Apocynaceae Shrub DIU-001 Aesthetic, Medicinal (antimicrobial)
2 Ficus Ficus benjamina Moraceae Tree DIU-002 Aesthetic, Environmental (air purification)
3 European buckthorn Rhamnus cathartica Rhamnaceae Shrub DIU-003 Medicinal (laxative), Economic
4 Shewra Streblus asper Moraceae Tree DIU-004 Medicinal (dental), Cultural
5 Common box Buxus sempervirens Buxaceae Shrub DIU-005 Aesthetic (topiary), Economic
6 Weeping debdaru Polyalthia longifolia Annonaceae Tree DIU-006 Cultural/Ritual, Aesthetic
7 Carpet grass Axonopus fissifolius Poaceae Herb DIU-007 Aesthetic (lawn), Environmental
8 Evergreen bamboo Bambusa pleioblastus pumilus Poaceae Grass DIU-008 Environmental, Economic
9 Umbrella grass Cyperus alternifolius Cyperaceae Herb DIU-009 Aesthetic, Medicinal
10 Broad-leaf lady palm Rhapis excelsa Arecaceae Palm DIU-010 Aesthetic (indoor)
11 Sunflower Helianthus annuus Asteraceae Herb DIU-011 Aesthetic, Nutritional, Medicinal
12 Chinese ixora Ixora chinensis Rubiaceae Shrub DIU-012 Aesthetic, Medicinal (anti-inflammatory)
13 Macarthur palm Ptychosperma macarthurii Arecaceae Palm DIU-013 Aesthetic (landscape)
14 German bamboo Pogonatherum crinitum Poaceae Grass DIU-014 Aesthetic, Environmental
15 Poison bulb / Spider lily Crinum asiaticum Amaryllidaceae Herb DIU-015 Medicinal (traditional), Cultural
16 Black matipo Pittosporum tenuifolium Pittosporaceae Shrub DIU-016 Aesthetic, Environmental
17 Areca palm Dypsis lutescens Arecaceae Palm DIU-017 Aesthetic, Environmental (air purifier)
18 Papuan fishtail palm Ptychosperma burretianum Arecaceae Palm DIU-018 Aesthetic
19 Leyland cypress Cupressocyparis leylandii Cupressaceae Tree DIU-019 Environmental (windbreak), Aesthetic
20 Ti plant Cordyline fruticosa Asparagaceae Shrub DIU-020 Cultural/Ritual, Aesthetic
21 Ming aralia Polyscias fruticosa Araliaceae Shrub DIU-021 Aesthetic, Medicinal
22 Pastureweed Cyathula prostrata Amaranthaceae Herb DIU-022 Medicinal (traditional)
23 Fishtail sword fern Nephrolepis falcata Nephrolepidaceae Fern DIU-023 Aesthetic (indoor)
24 Copperleaf plant Acalypha wilkesiana Euphorbiaceae Shrub DIU-024 Aesthetic, Medicinal
25 Jasmine Jasminum sp. Oleaceae Climber DIU-025 Aesthetic, Cultural, Medicinal (aromatherapy)
26 Dracaena Dracaena fragrans Asparagaceae Shrub DIU-026 Aesthetic, Environmental (air purifier)
27 Dwarf voodoo lily Typhonium roxburghii Araceae Herb DIU-027 Medicinal (traditional)
28 Thuja Thuja occidentalis Cupressaceae Tree DIU-028 Aesthetic, Medicinal
29 Euodia Euodia suaveolens Rutaceae Tree DIU-029 Medicinal (aromatic), Cultural
30 Japanese grass Microstegium vimineum Poaceae Grass DIU-030 Aesthetic, Environmental
31 Cycas Cycas revoluta Cycadaceae Cycad DIU-031 Aesthetic, Cultural
32 Palmyra palm Borassus flabellifer Arecaceae Palm DIU-032 Cultural, Nutritional, Economic
33 Euphorbia Chamaesyce heyneana Euphorbiaceae Herb DIU-033 Medicinal (traditional)
34 Yellow walking iris Trimezia steyermarkii Iridaceae Herb DIU-034 Aesthetic
35 Chamaedorea palm Chamaedorea elegans Arecaceae Palm DIU-035 Aesthetic (indoor)
36 Phyllanthus Cathetus myrtifolius Phyllanthaceae Shrub DIU-036 Medicinal, Environmental
37 Grass pea Lathyrus sativus Fabaceae Herb DIU-037 Nutritional, Environmental (nitrogen fixation)
38 Camellia Camellia japonica Theaceae Shrub DIU-038 Aesthetic, Cultural
39 Agave Agave americana Asparagaceae Herb DIU-039 Economic, Aesthetic, Medicinal
40 Wood fern Dryopteris filix-mas Dryopteridaceae Fern DIU-040 Aesthetic, Medicinal
41 Traveller’s tree Ravenala madagascariensis Strelitziaceae Tree DIU-041 Aesthetic, Cultural
42 Fukien tea Carmona retusa Boraginaceae Shrub DIU-042 Aesthetic (bonsai), Medicinal
43 Pauper’s tea Sageretia thea Rhamnaceae Shrub DIU-043 Aesthetic (bonsai), Cultural
44 Lucky bamboo Dracaena sanderiana Asparagaceae Herb DIU-044 Cultural/Ritual, Aesthetic
45 China rose Hibiscus rosa-sinensis Malvaceae Shrub DIU-045 Cultural (national flower), Medicinal, Aesthetic
46 False Ashoka Monoon longifolium Annonaceae Tree DIU-046 Aesthetic, Environmental
47 Oriental arborvitae Platycladus orientalis Cupressaceae Tree DIU-047 Aesthetic, Environmental, Medicinal
48 Japanese hawthorn Rhaphiolepis umbellata Rosaceae Shrub DIU-048 Aesthetic
49 Fire-flag Thalia geniculata Marantaceae Herb DIU-049 Aesthetic (aquatic), Environmental
50 Glossy privet Ligustrum lucidum Oleaceae Tree DIU-050 Aesthetic, Medicinal (traditional)
51 Alexander palm Ptychosperma elegans Arecaceae Palm DIU-051 Aesthetic (landscape)
52 Tea tree Melaleuca alternifolia Myrtaceae Tree DIU-052 Medicinal (antiseptic), Economic
53 Stone Gate palm Trachycarpus princeps Arecaceae Palm DIU-053 Aesthetic
54 Calla lily / Arum lily Zantedeschia aethiopica Araceae Herb DIU-054 Aesthetic, Cultural
55 Giant maidenhair fern Adiantum trapeziforme Pteridaceae Fern DIU-055 Aesthetic, Medicinal
56 Madagascar periwinkle Catharanthus roseus Apocynaceae Herb DIU-056 Medicinal (anticancer alkaloids), Aesthetic
57 Areca palm Chrysalidocarpus lutescens Arecaceae Palm DIU-057 Aesthetic, Environmental
58 Lemongrass Cymbopogon citratus Poaceae Grass DIU-058 Medicinal, Nutritional, Economic
59 Abyssinian banana Ensete ventricosum ‘Maurelii’ Musaceae Herb DIU-059 Aesthetic, Cultural
60 Red passion flower Passiflora coccinea Passifloraceae Climber DIU-060 Aesthetic, Medicinal
61 Violet crownshaft palm Pinanga speciosa Arecaceae Palm DIU-061 Aesthetic
62 Almond Prunus dulcis Rosaceae Tree DIU-062 Nutritional, Economic, Aesthetic
63 Spider lily Lycoris radiata Amaryllidaceae Herb DIU-063 Aesthetic, Cultural
64 Garden croton Codiaeum variegatum Euphorbiaceae Shrub DIU-064 Aesthetic, Medicinal
65 Bulblet fern Cystopteris bulbifera Dryopteridaceae Fern DIU-065 Aesthetic
66 False bird of paradise Heliconia rostrata Heliconiaceae Herb DIU-066 Aesthetic, Cultural
67 Heartleaf philodendron Philodendron hederaceum Araceae Climber DIU-067 Aesthetic (indoor), Environmental
68 Scarlet clock vine Thunbergia coccinea Acanthaceae Climber DIU-068 Aesthetic, Medicinal
Table 2. Importance Value Index (IVI) components and Use Value (UV) of the top 10 ornamental tree and shrub species on the DIU campus. RD = Relative Density; RF = Relative Frequency; RDom = Relative Dominance; max IVI = 300.
Table 2. Importance Value Index (IVI) components and Use Value (UV) of the top 10 ornamental tree and shrub species on the DIU campus. RD = Relative Density; RF = Relative Frequency; RDom = Relative Dominance; max IVI = 300.
Species RD (%) RF (%) RDom (%) IVI UV
Polyalthia longifolia 18.2 16.8 17.5 52.5 2.8
Ficus benjamina 15.4 14.2 14.8 44.4 2.4
Borassus flabellifer 12.8 13.5 13.2 39.5 2.1
Ravenala madagascariensis 11.6 11.2 10.8 33.6 1.9
Dypsis lutescens 10.9 10.4 10.2 31.5 1.7
Bambusa pleioblastus pumilus 9.7 9.8 9.5 29.0 1.5
Cordyline fruticosa 8.4 8.9 8.2 25.5 1.8
Ixora chinensis 7.6 7.5 7.2 22.3 2.0
Hibiscus rosa-sinensis 6.3 6.2 6.8 19.3 2.3
Cycas revoluta 5.8 5.9 5.6 17.3 1.6
Table 3. Ethnobotanical use categories, Informant Consensus Factor (ICF), and mean Use Value (UV) of ornamental plants on the DIU campus (n = 103 informants; 68 species).
Table 3. Ethnobotanical use categories, Informant Consensus Factor (ICF), and mean Use Value (UV) of ornamental plants on the DIU campus (n = 103 informants; 68 species).
Use Category No. Species % Species ICF Mean UV Key Uses Reported
Medicinal 28 41.2 0.62 1.92 Treating fever, skin diseases, digestive issues
Aesthetic/Ornamental 68 100.0 0.81 2.41 Campus beautification, landscape design
Cultural/Ritual 14 20.6 0.55 1.38 Religious ceremonies, festive decoration
Environmental 32 47.1 0.68 1.74 Air purification, shade, carbon sequestration
Economic 19 27.9 0.59 1.55 Timber, essential oils, market value
Nutritional 11 16.2 0.51 1.22 Edible fruits, seeds, leaves
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