Submitted:
01 July 2026
Posted:
02 July 2026
You are already at the latest version
Abstract
Keywords:
1. Introduction
2. Materials and Methods
2.1. Study Area and Occurrence Records
2.2. Environmental Variables Selection
2.3. Species Distribution Modeling
2.4. Model Evaluation and Variable Importance
2.5. Habitat Suitability Mapping
3. Results and Discussions
3.1. Environmental Variables Selection
3.2. Final Environmental Variables Influencing the Habitat Distribution of A. philippinensis
3.3. Predicted Habitat Suitability and Potential Distribution of A. philippinensis
3.4. Limitations of This Study
4. Conclusions
Supplementary Materials
Author Contributions
Funding
Data Availability Statement
Acknowledgments
Conflicts of Interest
Abbreviations
| Abbreviation | Full Term |
| A. philippinensis | Agathis philippinensis |
| AUC | Area Under the Receiver Operating Characteristic Curve |
| BIO | Bioclimatic variable |
| BIO1-BIO19 | WorldClim bioclimatic variables |
| BIO5 | Maximum Temperature of the Warmest Month |
| DEM | Digital Elevation Model |
| DENR | Department of Environment and Natural Resources |
| DorSU | Davao Oriental State University |
| ELEV | Elevation |
| EMBC | Eastern Mindanao Biodiversity Corridor |
| GBIF | Global Biodiversity Information Facility |
| GEF | Global Environment Facility |
| GIS | Geographic Information System |
| IUCN | International Union for Conservation of Nature |
| MaxEnt | Maximum Entropy |
| PRS | Philippine Reference System |
| ROC | Receiver Operating Characteristic |
| SDM | Species Distribution Modeling |
| SRTM | Shuttle Radar Topography Mission |
| MaxEnt | Maximum Entropy |
| UNDP | United Nations Development Programme |
| UTM | Universal Transverse Mercator |
| VIF | Variance Inflation Factor |
| VU | Vulnerable |
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| Code | BIOCLIM Variable | Description / Meaning |
| BIO1 | Annual Mean Temperature | Mean temperature across the year |
| BIO2 | Mean Diurnal Range | Mean of the monthly maximum temperature minus the minimum temperature |
| BIO3 | Isothermality | BIO2 divided by BIO7 × 100 |
| BIO4 | Temperature Seasonality | Variation in temperature across the year |
| BIO5 | Maximum Temperature of Warmest Month | Highest monthly maximum temperature |
| BIO6 | Minimum Temperature of Coldest Month | Lowest monthly minimum temperature |
| BIO7 | Temperature Annual Range | BIO5 minus BIO6 |
| BIO8 | Mean Temperature of Wettest Quarter | Mean temperature during the wettest 3-month period |
| BIO9 | Mean Temperature of Driest Quarter | Mean temperature during the driest 3-month period |
| BIO10 | Mean Temperature of Warmest Quarter | Mean temperature during the warmest 3-month period |
| BIO11 | Mean Temperature of Coldest Quarter | Mean temperature during the coldest 3-month period |
| BIO12 | Annual Precipitation | Total precipitation across the year |
| BIO13 | Precipitation of Wettest Month | Highest monthly precipitation |
| BIO14 | Precipitation of Driest Month | Lowest monthly precipitation |
| BIO15 | Precipitation Seasonality | Variation in precipitation across the year |
| BIO16 | Precipitation of the Wettest Quarter | Total precipitation during the wettest 3-month period |
| BIO17 | Precipitation of the Driest Quarter | Total precipitation during the driest 3-month period |
| BIO18 | Precipitation of Warmest Quarter | Total precipitation during the warmest 3-month period |
| BIO19 | Precipitation of Coldest Quarter | Total precipitation during the coldest 3-month period |
| Absolute Correlation Coefficient (|r|) | Interpretation | Action |
| < 0.70 | Acceptable correlation | Retain both variables |
| 0.70–0.79 | High correlation | Candidate for removal |
| 0.80–0.89 | Very high correlation | Remove one variable based on ecological relevance |
| ≥ 0.90 | Severe multicollinearity | Remove one variable immediately |
| AUC Value | Interpretation |
| 0.50-0.60 | Poor |
| 0.60-0.70 | Fair |
| 0.70-0.80 | Good |
| 0.80-0.90 | Very Good |
| >0.90 | Excellent |
| Suitability Class | Suitability Value |
| Unsuitable | 0.0-0.2 |
| Low | 0.2-0.4 |
| Moderate | 0.4-0.6 |
| High | 0.6-0.8 |
| Very High | 0.8-1.0 |
| Cluster | Variables | Representative |
| 1 | BIO4, BIO12, BIO13, BIO15, BIO16, BIO19 | BIO13 |
| 2 | BIO14, BIO17, BIO18 | BIO18 |
| 3 | BIO1, BIO5, BIO6, BIO8, BIO9, BIO10, BIO11 | BIO5 |
| 4 | BIO2, BIO3, BIO7 | BIO3 |
| 5 | ELEV | ELEV |
| Variable | Description | VIF |
| BIO5 | Maximum Temperature of the Warmest Month | 1.731 |
| ELEV | Elevation | 1.731 |
| Variable | Percent Contribution | Permutation Importance |
| ELEV | 90.5 | 93.2 |
| BIO5 | 9.5 | 6.2 |
| Class | Area (km²) | Percentage (%) |
| Unsuitable | 28,774.50 | 89.75 |
| Low | 1,801.02 | 5.62 |
| Moderate | 731.88 | 2.28 |
| High | 421.15 | 1.31 |
| Very High | 331.27 | 1.03 |
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